The Zopheridae family of beetles has grown considerably in recent years as the members of two other families have been included within its circumscription; these former families are the Monommatidae and the Colydiidae, which are now both included in the Zopheridae as subfamilies or (in the former case) even as tribe of subfamily Zopherinae. Some authors accept up to six subfamilies here, while others merge all except the Colydiinae into the Zopherinae.

Over 100 genera are in the redefined family, and hundreds of species worldwide. No common name is used for the new family, though some of the constituent subfamilies have their own, including the ironclad beetles, and the cylindrical bark beetles.

Ironclad female sjh
Ironclad beetle, Zopherus nodulosus haldemani
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: Polyphaga
Infraorder: Cucujiformia
Superfamily: Tenebrionoidea
Family: Zopheridae
Solier, 1834

Colydiinae - cylindrical bark beetles
Latometinae (sometimes in Zopheridae)
Monommatinae (sometimes in Zopheridae)
Pycnomerinae (sometimes in Zopheridae)
Usechinae (sometimes in Zopheridae)
Zopherinae - ironclad beetles

Zopheridae jewelry sjh
Zopheridae used as jewelry
Aulonium ruficorne (Olivier, 1790) (14425954610)
Aulonium ruficorne

See also


These 89 genera belong to the family Zopheridae:

  • Ablabus Broun, 1880 g
  • Acolobicus Sharp, 1894 i c g b
  • Acropis Burmeister, 1840 g
  • Anarmostes Pascoe, 1860 g
  • Anchomma Leconte, 1858 g
  • Antilissus Sharp, 1879 i c g
  • Aspathines Champion, 1888 i c g b
  • Aulonium Erichson, 1845 i c g b
  • Bitoma Herbst, 1793 i c g b
  • Catolaemus Sharp, 1894 g
  • Cerchanotus Erichson, 1845 g
  • Chorasus Sharp, 1882 g
  • Cicones Curtis, 1827 g
  • Colobicones Grouvelle, 1918 g
  • Colobicus Latreille, 1807 i c g b
  • Colydium Fabricius, 1792 i c g b
  • Colydodes Motschulsky, 1855 g
  • Corticus Germar, 1824 g
  • Coxelus Dejean, 1821 i c g b
  • Denophoelus Stephan, 1989 i c g b
  • Diodesma Latreille, 1829 g
  • Diplagia Reitter, 1882 g
  • Dryptops Broun, 1882 g
  • Endeitoma Sharp, 1894 i c g b
  • Endocoxelus Waterhouse, 1876 g
  • Endophloeus Dejean, 1834 g
  • Enhypnon Carter, 1919 g
  • Ethelema Pascoe, 1860 g
  • Eucicones Sharp, 1894 i c g b
  • Eudesma LeConte, 1863 i c g b
  • Glyphocryptus Sharp, 1885 g
  • Helioctamenus Schaufuss, 1882 g
  • Hyporhagus Thomson, 1860 i c g b
  • Hyporrhagus Thomson, 1860 g
  • Inscutomonomma Pic, 1951 g
  • Langelandia Aubé, 1842 g
  • Lasconotus Erichson, 1845 i c g b
  • Lemnis Harold, 1868 g
  • Lobogestoria Reitter, 1878 i c g b
  • Lyreus Aubé, 1861 i c g b
  • Mecedanum Erichson, 1845 g
  • Megataphrus Casey, 1890 i c g b
  • Microprius Fairmaire, 1868 i c g b
  • Microsicus Sharp, 1894 i c g
  • Monoedus Horn, 1882 i c g b
  • Monomma Klug, 1833 g
  • Myrmecoxenus g
  • Namunaria Reitter, 1882 i c g b
  • Nematidium Erichson, 1845 i c g b
  • Neotrichus Sharp, 1886 i c g
  • Noserinus Casey, 1907 g
  • Noserus g
  • Nosoderma Solier, 1841 g
  • Nosodomodes Reitter, 1922 g
  • Orthocerus g
  • Paha Dajoz, 1984 i c g b
  • Phellopsis LeConte, 1862 i c g b
  • Phloeodalis Erichson, 1845 g
  • Phloeodes LeConte, 1862 i c g b
  • Phloeonemus Erichson, 1845 i c g b
  • Phormesa Pascoe, 1863 g
  • Phreatus Pascoe, 1863 g
  • Priolomus Erichson, 1845 g
  • Pristoderus Hope, 1840 g
  • Prosteca Wollaston, 1860 g
  • Prostheca Wollaston, 1860 g
  • Pseudaulonium Reitter, 1877 g
  • Pseudendestes Lawrence, 1980 g
  • Pseudocorticus Hinton, 1935 i c g b
  • Pycnomerus Erichson, 1842 i c g b
  • Rechodes Erichson, 1845 g
  • Recyntus Broun, 1882 g
  • Rhagodera Mannerheim, 1843 i c g b
  • Rhagoderidea Wickham, 1914 g
  • Rhizonium Sharp, 1876 g
  • Sesaspis b
  • Spinhyporhagus Freude, 1984 i c g
  • Stephaniolus Ivie, Slipinsky and Wegrzynowicz, 2001 i c g
  • Synchita Hellwig, 1792 i c g b
  • Tarphiablabus Slipinski & Lawrence, 1997 g
  • Tarphiosoma Wollaston, 1862 g
  • Tarphius Erichson, 1845 g
  • Todimopsis Slipinski & Lawrence, 1997 g
  • Trigonophloeus Dajoz, 1980 g
  • Usechimorpha Blaisdell, 1929 i c g b
  • Usechus Motschulsky, 1845 i c g b
  • Vitiacus Broun, 1893 g
  • Xylolaemus Dejean, 1835 g
  • Zopherus Laporte, 1840 i c g b

Data sources: i = ITIS,[1] c = Catalogue of Life,[2] g = GBIF,[3] b = Bugguide.net[4]


  1. ^ "Zopheridae Report". Integrated Taxonomic Information System. Retrieved 2018-04-23.
  2. ^ "Browse Zopheridae". Catalogue of Life. Retrieved 2018-04-23.
  3. ^ "Zopheridae". GBIF. Retrieved 2018-04-23.
  4. ^ "Zopheridae Family Information". BugGuide.net. Retrieved 2018-04-23.

Aulonium is a genus of cylindrical bark beetles in the family Zopheridae. There are about five described species in Aulonium.


Bitoma is a genus of cylindrical bark beetles in the family Zopheridae. There are about 15 described species in Bitoma.


Colydiinae is a subfamily of beetles, commonly known as cylindrical bark beetles. They have been treated historically as a family Colydiidae, but have been moved into the Zopheridae, where they constitute the bulk of the diversity of the newly expanded family, with about 140 genera worldwide. They are diverse for example in the Australian region, from where about 35 genera are known; in Europe, though, only 20 genera are found and many of these only with few species.Little is known about the biology of these beetles. Most feed on fungi, others are carnivores and eat small arthropods such as bark beetles.


Colydium is a genus of cylindrical bark beetles in the family Zopheridae. There are about five described species in Colydium.


Eucicones is a genus of cylindrical bark beetles in the family Zopheridae. There is at least one described species in Eucicones, E. marginalis.


Lasconotus is a genus of cylindrical bark beetles in the family Zopheridae. There are at least 20 described species in Lasconotus.


Megataphrus is a genus of cylindrical bark beetles in the family Zopheridae. There are at least three described species in Megataphrus.


Monommatini is a tribe of beetles known as monommatid beetles. They are in the ironclad beetle family, Zopheridae. There are about 15 genera and 300 described species in Monommatini. They are found worldwide, with the greatest diversity in Madagascar. They are commonly associated with rotting plant matter such as the dry rotting cambium of trees.


Namunaria is a genus of cylindrical bark beetles in the family Zopheridae. There are at least two described species in Namunaria.


"Phloeodes" redirects here. This can also be a misspelling of Phlaeodes, nowadays a junior synonym of the tortrix moth genus Epinotia.

The genus Nosoderma includes 28 species of ironclad beetles from the Americas, including some common and widely distributed species placed in the former genus Phloeodes.

Nosoderma diabolicum

Nosoderma diabolicum (formerly Phloeodes diabolicus) is a beetle of the Family Zopheridae.

Nosoderma plicatum

Nosoderma plicatum (formerly Phloeodes plicatus) is a beetle of the Family Zopheridae.


Phloeonemus is a genus of cylindrical bark beetles in the family Zopheridae. There are at least two described species in Phloeonemus.


Pycnomerus is a genus of ironclad beetles in the family Zopheridae. There are at least 20 described species in Pycnomerus.


Synchitini is a tribe of cylindrical bark beetles in the family Zopheridae. There are about 19 genera and at least 40 described species in Synchitini.


The Tenebrionoidea are a very large and diverse superfamily of beetles.

It generally corresponds to the Heteromera of earlier authors.

It contains these families:

Aderidae Winkler 1927 (ant-like leaf beetles)

Anthicidae Latreille 1819 (ant-like flower beetles)

Archeocrypticidae Kaszab 1964

Boridae C. G. Thomson 1859

Chalcodryidae Watt 1974

Ciidae Leach 1819 (minute tree-fungus beetles) (= Cisidae)

Melandryidae Leach 1815 (false darkling beetles)

Meloidae Gyllenhal 1810 (blister beetles)

Mordellidae Latreille 1802 (tumbling flower beetles)

Mycetophagidae Leach 1815 (hairy fungus beetles)

Mycteridae Blanchard 1845

Oedemeridae Latreille 1810 (false blister beetles)

Perimylopidae St. George 1939

Prostomidae C. G. Thomson 1859

Pterogeniidae Crowson 1953

Pyrochroidae Latreille 1807 (fire-colored beetles, etc.)

Pythidae Solier 1834

Ripiphoridae Gemminger and Harold 1870 (wedge-shaped beetles) (= Rhipiphoridae)

Salpingidae Leach 1815 (narrow-waisted bark beetles, etc.)

Scraptiidae Mulsant 1856 (false flower beetles)

Stenotrachelidae C. G. Thomson 1859 (false long-horned beetles) (= Cephaloidae)

Synchroidae Lacordaire 1859

Tenebrionidae Latreille 1802 (darkling beetles)

Tetratomidae Billberg 1820

Trachelostenidae Lacordaire 1859

Trictenotomidae Blanchard 1845

Ulodidae Pascoe 1869

Zopheridae Solier 1834 (ironclad beetles, cylindrical bark beetles, etc.)

Xylolaemus sakhnovi

Xylolaemus sakhnovi is an extinct species of cylindrical bark beetle in the family Zopheridae. The species is solely known from the Middle Eocene Baltic amber deposits in the Baltic Sea region of Europe. The genus Xylolaemus contains a total of six extant species distributed from western Europe through the Canary Islands and North Africa to India. The species is the first in the genus to be described from a fossil specimen.


Zopherinae is a subfamily of beetles, commonly known as ironclad beetles. Together with the subfamily Usechinae, they have been treated historically as a family, but have recently been joined by several additional taxa, making the Zopheridae a much larger composite family, and the Zopherinae are now only a small component within it, consisting of seven genera in the tribe Zopherini and one genus, Phellopsis in its own tribe (Phellopsini).

These beetles are apparently fungivores and associated with rotting wood, and as the common name implies, have one of the hardest of all arthropod exoskeletons; in some species, it is almost impossible to drive an insect pin through their bodies without using a small drill to make a hole first.

When disturbed, ironclad beetles play dead.

Some species in the genus Zopherus in Mexico are decorated with costume jewelry glued to their bodies, and sold as living brooches, known as ma'kech.


Zopherus is a genus of beetles comprising 19 species. They live in the Americas and are adapted to wood-boring. Some species are used as living brooches.

Extant Coleoptera families


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