The Triassic ( /traɪˈæsɪk/) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.9 million years ago (Mya), to the beginning of the Jurassic Period 201.3 Mya. The Triassic is the first period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events.
Triassic began in the wake of the Permian–Triassic extinction event, which left the Earth's biosphere impoverished; it was well into the middle of the Triassic before life recovered its former diversity. Therapsids and archosaurs were the chief terrestrial vertebrates during this time. A specialized subgroup of archosaurs, called dinosaurs, first appeared in the Late Triassic but did not become dominant until the succeeding Jurassic Period.
The first true mammals, themselves a specialized subgroup of therapsids, also evolved during this period, as well as the first flying vertebrates, the pterosaurs, who, like the dinosaurs, were a specialized subgroup of archosaurs. The vast supercontinent of Pangaea existed until the mid-Triassic, after which it began to gradually rift into two separate landmasses, Laurasia to the north and Gondwana to the south.
The global climate during the Triassic was mostly hot and dry, with deserts spanning much of Pangaea's interior. However, the climate shifted and became more humid as Pangaea began to drift apart. The end of the period was marked by yet another major mass extinction, the Triassic–Jurassic extinction event, that wiped out many groups and allowed dinosaurs to assume dominance in the Jurassic.
The Triassic was named in 1834 by Friedrich von Alberti, after the three distinct rock layers (tri meaning "three") that are found throughout Germany and northwestern Europe—red beds, capped by marine limestone, followed by a series of terrestrial mud- and sandstones—called the "Trias".
251.902–201.3 million years ago
|Mean atmospheric O
2 content over period duration
|c. 16 vol %|
(80 % of modern level)
|Mean atmospheric CO
2 content over period duration
|c. 1750 ppm|
(6 times pre-industrial level)
|Mean surface temperature over period duration||c. 17 °C|
(3 °C above modern level)
The Triassic is usually separated into Early, Middle, and Late Triassic Epochs, and the corresponding rocks are referred to as Lower, Middle, or Upper Triassic. The faunal stages from the youngest to oldest are:
|Upper/Late Triassic (Tr3)|
|Rhaetian||(208.5 – 201.3 Mya)|
|Norian||(227 – 208.5 Mya)|
|Carnian||(237– 227 Mya)|
|Middle Triassic (Tr2)|
|Ladinian||(242 – 237 Mya)|
|Anisian||(247.2 – 242 Mya)|
|Lower/Early Triassic (Scythian)|
|Olenekian||(251.2 – 247.2 Mya)|
|Induan||(251.902 – 251.2 Mya)|
During the Triassic, almost all the Earth's land mass was concentrated into a single supercontinent centered more or less on the equator and spanning from pole to pole, called Pangaea ("all the land"). From the east, along the equator, the Tethys sea penetrated Pangaea, causing the Paleo-Tethys Ocean to be closed.
Later in the mid-Triassic a similar sea penetrated along the equator from the west. The remaining shores were surrounded by the world-ocean known as Panthalassa ("all the sea"). All the deep-ocean sediments laid down during the Triassic have disappeared through subduction of oceanic plates; thus, very little is known of the Triassic open ocean.
The supercontinent Pangaea was rifting during the Triassic—especially late in that period—but had not yet separated. The first nonmarine sediments in the rift that marks the initial break-up of Pangaea, which separated New Jersey from Morocco, are of Late Triassic age; in the U.S., these thick sediments comprise the Newark Group.
Because a super-continental mass has less shoreline compared to one broken up, Triassic marine deposits are globally relatively rare, despite their prominence in Western Europe, where the Triassic was first studied. In North America, for example, marine deposits are limited to a few exposures in the west. Thus Triassic stratigraphy is mostly based on organisms that lived in lagoons and hypersaline environments, such as Estheria crustaceans.
At the beginning of the Mesozoic Era, Africa was joined with Earth's other continents in Pangaea. Africa shared the supercontinent's relatively uniform fauna which was dominated by theropods, prosauropods and primitive ornithischians by the close of the Triassic period. Late Triassic fossils are found throughout Africa, but are more common in the south than north. The time boundary separating the Permian and Triassic marks the advent of an extinction event with global impact, although African strata from this time period have not been thoroughly studied.
During the Triassic peneplains are thought to have formed in what is now Norway and southern Sweden. Remnants of this peneplain can be traced as a tilted summit accordance in the Swedish West Coast. In northern Norway Triassic peneplains may have been buried in sediments to be then re-exposed as coastal plains called strandflats. Dating of illite clay from a strandflat of Bømlo, southern Norway, have shown that landscape there became weathered in Late Triassic times (c. 210 million years ago) with the landscape likely also being shaped during that time.
At Paleorrota geopark, located in Rio Grande do Sul, Brazil, the Santa Maria Formation and Caturrita Formations are exposed. In these formations, one of the earliest dinosaurs, Staurikosaurus, as well as the mammal ancestors Brasilitherium and Brasilodon have been discovered.
The Triassic continental interior climate was generally hot and dry, so that typical deposits are red bed sandstones and evaporites. There is no evidence of glaciation at or near either pole; in fact, the polar regions were apparently moist and temperate, providing a climate suitable for forests and vertebrates, including reptiles. Pangaea's large size limited the moderating effect of the global ocean; its continental climate was highly seasonal, with very hot summers and cold winters. The strong contrast between the Pangea supercontinent and the global ocean triggered intense cross-equatorial monsoons.
The Triassic may have mostly been a dry period, but evidence exists that it was punctuated by several episodes of increased rainfall in tropical and subtropical latitudes of the Tethys Sea and its surrounding land. Sediments and fossils suggestive of a more humid climate are known from the Anisian to Ladinian of the Tethysian domain, and from the Carnian and Rhaetian of a larger area that includes also the Boreal domain (e.g., Svalbard Islands), the North American continent, the South China block and Argentina.
The best studied of such episodes of humid climate, and probably the most intense and widespread, was the Carnian Pluvial Event.
Three categories of organisms can be distinguished in the Triassic record: survivors from the Permian–Triassic extinction event, new groups which flourished briefly, and other new groups which went on to dominate the Mesozoic Era.
On land, the surviving vascular plants included the lycophytes, the dominant cycadophytes, ginkgophyta (represented in modern times by Ginkgo biloba), ferns, horsetails and glossopterids. The spermatophytes, or seed plants, came to dominate the terrestrial flora: in the northern hemisphere, conifers, ferns and bennettitales flourished. Glossopteris (a seed fern) was the dominant southern hemisphere tree during the Early Triassic period.
In marine environments, new modern types of corals appeared in the Early Triassic, forming small patches of reefs of modest extent compared to the great reef systems of Devonian or modern times. Serpulids appeared in the Middle Triassic. Microconchids were abundant. The shelled cephalopods called ammonites recovered, diversifying from a single line that survived the Permian extinction.
The fish fauna was remarkably uniform, suggesting that very few families survived the Permian extinction. There were also many types of marine reptiles. These included the Sauropterygia, which featured pachypleurosaurus and nothosaurs (both common during the Middle Triassic, especially in the Tethys region), placodonts, and the first plesiosaurs. The first of the lizardlike Thalattosauria (askeptosaurs) and the highly successful ichthyosaurs, which appeared in Early Triassic seas soon diversified, and some eventually developed to huge size during the Late Triassic. Subequatorial saurichthyids have also been described in Early Triassic strata.
The Permian–Triassic extinction devastated terrestrial life. Biodiversity rebounded as the surviving species repopulated empty terrain, but these were short-lived. Diverse communities with complex food-web structures took 30 million years to reestablish.
Temnospondyl amphibians were among those groups that survived the Permian–Triassic extinction; some lineages (e.g. trematosaurs) flourished briefly in the Early Triassic, while others (e.g. capitosaurs) remained successful throughout the whole period, or only came to prominence in the Late Triassic (e.g. plagiosaurs, metoposaurs). As for other amphibians, the first Lissamphibia, progenitors of first frogs, are known from the Early Triassic, but the group as a whole did not become common until the Jurassic, when the temnospondyls had become very rare. Other survivors the Chroniosuchia and Embolomeri were more closely related to amniotes than temnospondyls. Those became extinct after some million years.
Most of the Reptiliomorpha, stem-amniotes that gave rise to the amniotes, disappeared in Triassic, but two water-dwelling groups survived; Embolomeri that only survived into the early part of the period, and the Chroniosuchia, which survived until the end of Triassic.
Archosauromorph reptiles, especially archosaurs, progressively replaced the synapsids that had dominated the previous Permian period. The Cynognathus was the characteristic top predator in earlier Triassic (Olenekian and Anisian) on Gondwana. Both kannemeyeriid dicynodonts and gomphodont cynodonts remained important herbivores during much of the period, and ecteniniids played a role as large-sized, cursorial predators in the Late Triassic. During the Carnian (early part of the Late Triassic), some advanced cynodonts gave rise to the first mammals. At the same time the Ornithodira, which until then had been small and insignificant, evolved into pterosaurs and a variety of dinosaurs. The Crurotarsi were the other important archosaur clade, and during the Late Triassic these also reached the height of their diversity, with various groups including the phytosaurs, aetosaurs, several distinct lineages of Rauisuchia, and the first crocodylians (the Sphenosuchia). Meanwhile, the stocky herbivorous rhynchosaurs and the small to medium-sized insectivorous or piscivorous Prolacertiformes were important basal archosauromorph groups throughout most of the Triassic.
Among other reptiles, the earliest turtles, like Proganochelys and Proterochersis, appeared during the Norian Age (Stage) of the Late Triassic Period. The Lepidosauromorpha, specifically the Sphenodontia, are first found in the fossil record of the earlier Carnian Age. The Procolophonidae were an important group of small lizard-like herbivores.
During the Triassic, archosaurs displaced therapsids as the dominant amniotes. This "Triassic Takeover" may have contributed to the evolution of mammals by forcing the surviving therapsids and their mammaliaform successors to live as small, mainly nocturnal insectivores. Nocturnal life may have forced the mammaliaforms to develop fur and a higher metabolic rate.
The Monte San Giorgio lagerstätte, now in the Lake Lugano region of northern Italy and Switzerland, was in Triassic times a lagoon behind reefs with an anoxic bottom layer, so there were no scavengers and little turbulence to disturb fossilization, a situation that can be compared to the better-known Jurassic Solnhofen Limestone lagerstätte.
The remains of fish and various marine reptiles (including the common pachypleurosaur Neusticosaurus, and the bizarre long-necked archosauromorph Tanystropheus), along with some terrestrial forms like Ticinosuchus and Macrocnemus, have been recovered from this locality. All these fossils date from the Anisian/Ladinian transition (about 237 million years ago).
The Triassic period ended with a mass extinction, which was particularly severe in the oceans; the conodonts disappeared, as did all the marine reptiles except ichthyosaurs and plesiosaurs. Invertebrates like brachiopods, gastropods, and molluscs were severely affected. In the oceans, 22% of marine families and possibly about half of marine genera went missing.
Though the end-Triassic extinction event was not equally devastating in all terrestrial ecosystems, several important clades of crurotarsans (large archosaurian reptiles previously grouped together as the thecodonts) disappeared, as did most of the large labyrinthodont amphibians, groups of small reptiles, and some synapsids (except for the proto-mammals). Some of the early, primitive dinosaurs also became extinct, but more adaptive ones survived to evolve into the Jurassic. Surviving plants that went on to dominate the Mesozoic world included modern conifers and cycadeoids.
The cause of the Late Triassic extinction is uncertain. It was accompanied by huge volcanic eruptions that occurred as the supercontinent Pangaea began to break apart about 202 to 191 million years ago (40Ar/39Ar dates), forming the Central Atlantic Magmatic Province (CAMP), one of the largest known inland volcanic events since the planet had first cooled and stabilized. Other possible but less likely causes for the extinction events include global cooling or even a bolide impact, for which an impact crater containing Manicouagan Reservoir in Quebec, Canada, has been singled out. However, the Manicouagan impact melt has been dated to 214±1 Mya. The date of the Triassic-Jurassic boundary has also been more accurately fixed recently, at 201.3 Mya. Both dates are gaining accuracy by using more accurate forms of radiometric dating, in particular the decay of uranium to lead in zircons formed at time of the impact. So, the evidence suggests the Manicouagan impact preceded the end of the Triassic by approximately 10±2 Ma. It could not therefore be the immediate cause of the observed mass extinction.
The number of Late Triassic extinctions is disputed. Some studies suggest that there are at least two periods of extinction towards the end of the Triassic, separated by 12 to 17 million years. But arguing against this is a recent study of North American faunas. In the Petrified Forest of northeast Arizona there is a unique sequence of late Carnian-early Norian terrestrial sediments. An analysis in 2002 found no significant change in the paleoenvironment. Phytosaurs, the most common fossils there, experienced a change-over only at the genus level, and the number of species remained the same. Some aetosaurs, the next most common tetrapods, and early dinosaurs, passed through unchanged. However, both phytosaurs and aetosaurs were among the groups of archosaur reptiles completely wiped out by the end-Triassic extinction event.
It seems likely then that there was some sort of end-Carnian extinction, when several herbivorous archosauromorph groups died out, while the large herbivorous therapsids—the kannemeyeriid dicynodonts and the traversodont cynodonts—were much reduced in the northern half of Pangaea (Laurasia).
These extinctions within the Triassic and at its end allowed the dinosaurs to expand into many niches that had become unoccupied. Dinosaurs became increasingly dominant, abundant and diverse, and remained that way for the next 150 million years. The true "Age of Dinosaurs" is during the following Jurassic and Cretaceous periods, rather than the Triassic.
In the geologic timescale, the Anisian is the lower stage or earliest age of the Middle Triassic series or epoch and lasted from 247.2 million years ago until 242 million years ago. The Anisian age succeeds the Olenekian age (part of the Lower Triassic epoch) and precedes the Ladinian age.Archosaur
Archosaurs are a group of diapsid amniotes whose living representatives consist of birds and crocodilians. This group also includes all extinct dinosaurs, extinct crocodilian relatives, and pterosaurs. Archosauria, the archosaur clade, is a crown group that includes the most recent common ancestor of living birds and crocodilians. It includes two main clades: Pseudosuchia, which includes crocodilians and their extinct relatives, and Avemetatarsalia, which includes birds and their extinct relatives (such as non-avian dinosaurs and pterosaurs).Avemetatarsalia
Avemetatarsalia (meaning "bird metatarsals") is a clade name established by British palaeontologist Michael Benton in 1999 for all crown group archosaurs that are closer to birds than to crocodiles. It includes a similarly defined subgroup, Ornithodira. An alternate name is Pan-Aves, or "all birds", in reference to its definition containing all animals, living or extinct, which are more closely related to birds than to crocodiles.
Members of this group include the Dinosauromorpha, Pterosauromorpha, the genus Scleromochlus, and Aphanosauria. Dinosauromorpha contains more basal forms, including Lagerpeton and Marasuchus, as well as more derived forms, including dinosaurs. Birds belong to the dinosaurs as members of the theropods. Pterosauromorpha contains Pterosauria, which were the first vertebrates capable of true flight. Aphanosauria is a Triassic group of gracile carnivorous quadrupeds which was recognized in 2017.Dinosauriformes
Dinosauriformes is a clade of archosaurian reptiles that include the dinosaurs and their most immediate relatives. All dinosauriformes are distinguished by several features, such as shortened forelimbs and a partially to fully perforated acetabulum, the hole in the hip socket traditionally used to define dinosaurs. The oldest known member is Asilisaurus, dating to about 245 million years ago in the Anisian age of the middle Triassic period.Dinosauromorpha
Dinosauromorpha is a clade of archosaurs that includes the clade Dinosauria (dinosaurs), and all animals more closely related to dinosaurs than to pterosaurs. Birds are the only surviving dinosauromorphs.Early Triassic
The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between 251.902 Ma and 247.2 Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic, which is a unit in chronostratigraphy. The Early Triassic is the oldest epoch of the Mesozoic Era and is divided into the Induan and Olenekian ages.
The Lower Triassic series is coeval with the Scythian stage, which is today not included in the official timescales but can be found in older literature. In Europe, most of the Lower Triassic is composed of Buntsandstein, a lithostratigraphic unit of continental red beds.Jurassic
The Jurassic (; from Jura Mountains) was a geologic period and system that spanned 56 million years from the end of the Triassic Period 201.3 million years ago (Mya) to the beginning of the Cretaceous Period 145 Mya. The Jurassic constitutes the middle period of the Mesozoic Era, also known as the Age of Reptiles. The start of the period was marked by the major Triassic–Jurassic extinction event. Two other extinction events occurred during the period: the Pliensbachian-Toarcian extinction in the Early Jurassic, and the Tithonian event at the end; however, neither event ranks among the "Big Five" mass extinctions.
The Jurassic period is divided into three epochs: Early, Middle, and Late. Similarly, in stratigraphy, the Jurassic is divided into the Lower Jurassic, Middle Jurassic, and Upper Jurassic series of rock formations.
The Jurassic is named after the Jura Mountains within the European Alps, where limestone strata from the period were first identified.
By the beginning of the Jurassic, the supercontinent Pangaea had begun rifting into two landmasses: Laurasia to the north, and Gondwana to the south. This created more coastlines and shifted the continental climate from dry to humid, and many of the arid deserts of the Triassic were replaced by lush rainforests.
On land, the fauna transitioned from the Triassic fauna, dominated by both dinosauromorph and crocodylomorph archosaurs, to one dominated by dinosaurs alone. The first birds also appeared during the Jurassic, having evolved from a branch of theropod dinosaurs. Other major events include the appearance of the earliest lizards, and the evolution of therian mammals, including primitive placentals. Crocodilians made the transition from a terrestrial to an aquatic mode of life. The oceans were inhabited by marine reptiles such as ichthyosaurs and plesiosaurs, while pterosaurs were the dominant flying vertebrates.Late Triassic
The Late Triassic is the third and final of three epochs of the Triassic Period in the geologic timescale. The Triassic-Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. The corresponding series is known as the Upper Triassic. In Europe the epoch was called the Keuper, after a German lithostratigraphic group (a sequence of rock strata) that has a roughly corresponding age. The Late Triassic spans the time between 237 Ma and 201.3 Ma (million years ago). The Late Triassic is divided into the Carnian, Norian and Rhaetian ages.
Many of the first dinosaurs evolved during the Late Triassic, including Plateosaurus, Coelophysis, and Eoraptor.
The extinction event that began during the Late Triassic resulted in the disappearance of about 76% of all terrestrial and marine life species, as well as almost 20% of taxonomic families. Although the Late Triassic Epoch did not prove to be as destructive as the preceding Permian Period, which took place approximately 50 million years earlier and destroyed about 70% of land species, 57% of insect families as well as 95% of marine life, it resulted in great decreased in population sizes of many living organism populations.
Specifically, the Late Triassic had negative effects on the conodonts and ammonoid groups. These groups once served as vital index fossils, which made it possible to identify feasible life span to multiple strata of the Triassic strata. These groups were severely affected during the epoch, and became extinct soon after. Despite the large populations that withered away with the coming of the Late Triassic, many families, such as the pterosaurs, crocodiles, mammals and fish were very minimally affected. However, such families as the bivalves, gastropods, marine reptiles and brachiopods were greatly affected and many species became extinct during this time.Lepidosauromorpha
Lepidosauromorpha is a group of reptiles comprising all diapsids closer to lizards than to archosaurs (which include crocodiles and birds). The only living sub-group is the Lepidosauria: extant lizards, snakes, amphisbaenians and tuataras.
Lepidosauromorpha are distinguishable from Archosauromorphs (archosaurs) by their primitive sprawling gait, which allows for the same sinusoidal trunk and tail movement seen in fish, the sliding "joint" between the coracoids and the sternum (for a longer stride), and their pleurodont dentition. In contrast, Archosauromorphs possess a parasagittal gait, a reduction in their dermal girdle, a reduction and/or loss of the sternum, and a more thecodont dentition.
Lepidosauromorpha have retained cold blood because of their low-energy sprawling stance.Mesozoic
The Mesozoic Era ( or ) is an interval of geological time from about 252 to 66 million years ago. It is also called the Age of Reptiles, a phrase introduced by the 19th century paleontologist Gideon Mantell who viewed it as dominated by diapsids such as Iguanodon, Megalosaurus, Plesiosaurus and Pterodactylus. To paleobotanists, this Era is also called the Age of Conifers.Mesozoic means "middle life", deriving from the Greek prefix meso-/μεσο- for "between" and zōon/ζῷον meaning "animal" or "living being". The name "Mesozoic" was proposed in 1840 by the British geologist John Phillips (1800–1874). It is one of three geologic eras of the Phanerozoic Eon, preceded by the Paleozoic ("ancient life") and succeeded by the Cenozoic ("new life"). The era is subdivided into three major periods: the Triassic, Jurassic, and Cretaceous, which are further subdivided into a number of epochs and stages.
The era began in the wake of the Permian–Triassic extinction event, the largest well-documented mass extinction in Earth's history, and ended with the Cretaceous–Paleogene extinction event, another mass extinction whose victims included the non-avian dinosaurs. The Mesozoic was a time of significant tectonic, climate and evolutionary activity. The era witnessed the gradual rifting of the supercontinent Pangaea into separate landmasses that would move into their current positions during the next era. The climate of the Mesozoic was varied, alternating between warming and cooling periods. Overall, however, the Earth was hotter than it is today. Dinosaurs appeared in the Late Triassic and became the dominant terrestrial vertebrates early in the Jurassic, occupying this position for about 135 million years until their demise at the end of the Cretaceous. Birds first appeared in the Jurassic, having evolved from a branch of theropod dinosaurs. The first mammals also appeared during the Mesozoic, but would remain small—less than 15 kg (33 lb)—until the Cenozoic.Middle Triassic
In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided. It spans the time between 247.2 Ma and 237 Ma (million years ago). The Middle Triassic is divided into the Anisian and Ladinian ages or stages.
Formerly the middle series in the Triassic was also known as Muschelkalk. This name is now only used for a specific unit of rock strata with approximately Middle Triassic age, found in western Europe.
During this time there were no flowering plants, but instead there were ferns and mosses. Small dinosaurs began to appear like Nyasasaurus and the ichnogenus Iranosauripus.Permian
The Permian is a geologic period and system which spans 47 million years from the end of the Carboniferous Period 298.9 million years ago (Mya), to the beginning of the Triassic period 251.902 Mya. It is the last period of the Paleozoic era; the following Triassic period belongs to the Mesozoic era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the city of Perm.
The Permian witnessed the diversification of the early amniotes into the ancestral groups of the mammals, turtles, lepidosaurs, and archosaurs. The world at the time was dominated by two continents known as Pangaea and Siberia, surrounded by a global ocean called Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, who could better cope with these drier conditions, rose to dominance in place of their amphibian ancestors.
The Permian (along with the Paleozoic) ended with the Permian–Triassic extinction event, the largest mass extinction in Earth's history, in which nearly 96% of marine species and 70% of terrestrial species died out. It would take well into the Triassic for life to recover from this catastrophe. Recovery from the Permian–Triassic extinction event was protracted; on land, ecosystems took 30 million years to recover.Permian–Triassic extinction event
The Permian–Triassic (P–Tr or P–T) extinction event, colloquially known as the Great Dying, the End-Permian Extinction or the Great Permian Extinction, occurred about 252 Ma (million years) ago, forming the boundary between the Permian and Triassic geologic periods, as well as between the Paleozoic and Mesozoic eras. It is the Earth's most severe known extinction event, with up to 96% of all marine species and 70% of terrestrial vertebrate species becoming extinct. It is the only known mass extinction of insects. Some 57% of all biological families and 83% of all genera became extinct. Because so much biodiversity was lost, the recovery of land-dwelling life took significantly longer than after any other extinction event, possibly up to 10 million years. Studies in Bear Lake County near Paris, Idaho showed a relatively quick rebound in a localized marine ecosystem, taking around 2 million years to recover, suggesting that the impact of the extinction may have been felt less severely in some areas than others.
There is evidence for one to three distinct pulses, or phases, of extinction. Suggested mechanisms for the latter include one or more large meteor impact events, massive volcanism such as that of the Siberian Traps, and the ensuing coal or gas fires and explosions, and a runaway greenhouse effect triggered by sudden release of methane from the sea floor due to methane clathrate dissociation according to the clathrate gun hypothesis or methane-producing microbes known as methanogens. Possible contributing gradual changes include sea-level change, increasing anoxia, increasing aridity, and a shift in ocean circulation driven by climate change.Rhaetian
See Raetians for the Alpine people of antiquity. See Raetian language for their language.The Rhaetian is, in geochronology, the latest age of the Triassic period or in chronostratigraphy the uppermost stage of the Triassic system. It lasted from 208.5 to 201.3 million years ago. It was preceded by the Norian and succeeded by the Hettangian (the lowermost stage or earliest age of the Jurassic).In this age, Pangaea began to break up, though the Atlantic Ocean was not yet formed.Sauropterygia
Sauropterygia ("lizard flippers") is an extinct, diverse taxon of aquatic reptiles that developed from terrestrial ancestors soon after the end-Permian extinction and flourished during the Triassic before all except for the Plesiosauria became extinct at the end of that era. The plesiosaurs would continue to diversify till the end of the Mesozoic. Sauropterygians are united by a radical adaptation of their pectoral girdle, adapted to support powerful flipper strokes. Some later sauropterygians, such as the pliosaurs, developed a similar mechanism in their pelvis.Therapsid
Therapsida is a group of synapsids that includes mammals and their ancestors. Many of the traits today seen as unique to mammals had their origin within early therapsids, including having their four limbs extend vertically beneath the body, as opposed to the sprawling posture of reptiles. The earliest fossil attributed to Therapsida is Tetraceratops insignis from the Lower Permian.Therapsids evolved from "pelycosaurs", specifically within the Sphenacodontia, more than 275 million years ago. They replaced the "pelycosaurs" as the dominant large land animals in the Middle Permian and were largely replaced, in turn, by the archosauromorphs in the Triassic, although one group of therapsids, the kannemeyeriiforms, remained diverse in the Late Triassic.
The therapsids included the cynodonts, the group that gave rise to mammals in the Late Triassic around 225 million years ago. Of the non-mammalian therapsids, only cynodonts survived the Triassic–Jurassic extinction event. The last of the non-mammalian therapsids, the tritylodontid cynodonts, became extinct in the Early Cretaceous, approximately 100 million years ago.Triassic–Jurassic extinction event
The Triassic–Jurassic extinction event marks the boundary between the Triassic and Jurassic periods, 201.3 million years ago, and is one of the major extinction events of the Phanerozoic eon, profoundly affecting life on land and in the oceans. In the seas, a whole class (conodonts) and 23-34% of marine genera disappeared. On land, all archosaurs other than crocodylomorphs (Sphenosuchia and Crocodyliformes) and Avemetatarsalia (pterosaurs and dinosaurs), some remaining therapsids, and many of the large amphibians became extinct.