The Tephritoidea are a superfamily of flies. The following families are included:

The Tachiniscinae, formerly ranked as the family Tachiniscidae, are now included in the Tephritidae.

Physiphora alceae female
Female Physiphora alceae
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Diptera
Subsection: Acalyptratae
Superfamily: Tephritoidea

see text


External links

  • "Tephritoidea". Atlas of Living Australia.

The Acalyptratae or Acalyptrata are a subsection of the Schizophora, which are a section of the order Diptera, the "true flies". In various contexts the Acalyptratae also are referred to informally as the acalyptrate muscoids, or acalyptrates, as opposed to the Calyptratae. All forms of the name refer to the lack of calypters in the members of this subsection of flies. An alternative name, Acalypterae is current, though in minority usage. It was first used by Justin Pierre Marie Macquart in 1835 for a section of his tribe Muscides; he used it to refer to all acalyptrates plus scathophagids and phorids, but excluding Conopidae.

The confusing forms of the names stem from their first usage; Acalyptratae and Acalyptrata actually are adjectival forms in New Latin. They were coined in the mid 19th century in contexts such as "Muscae Calyptratae and Acalyptratae" and "Diptera Acalyptrata", and the forms stuck.The Acalyptratae are a large assemblage, exhibiting very diverse habits, with one notable and perhaps surprising exception: no known acalyptrates are obligate blood-feeders (hematophagous), though blood feeding at various stages of the life history is common throughout other Dipteran sections.


Aciuroides is a genus of ulidiid or picture-winged fly in the family Ulidiidae.


Acrosticta is a genus of picture-winged flies in the family Ulidiidae.


Automola is a genus of flies in the family Richardiidae. There are at least three described species in Automola.

Automola atomaria

Automola atomaria is a species of fly in the genus Automola of the family Richardiidae. A. atomaria is found in Central America.

Cheese fly

The cheese fly (Piophila casei) is a species of fly known for infesting human foodstuffs. The larvae of this fly are known as cheese skippers due to their ability to launch themselves several inches into the air when alarmed. When consumed, the larvae can survive in the intestine, causing enteric myiasis.


Idana is a genus of flies in the family Ulidiidae. There is at least one described species in Idana, I. marginata.


The Brachyceran infraorder Muscomorpha is a large and diverse group of flies, containing the bulk of the Brachycera, and, most of the known flies. It includes a number of the most familiar flies, such as the housefly, the fruit fly, and the blow fly. The antennae are short, usually three-segmented, with a dorsal arista. Their bodies are often highly setose, and the pattern of setae is often taxonomically important.

The larvae of muscomorphs (in the sense the name is used here; see below) have reduced head capsules, and the pupae are formed inside the exoskeleton of the last larval instar; exit from this puparium is by a circular line of weakness, and this pupal type is called "cyclorrhaphous"; this feature gives this group of flies their traditional name, Cyclorrhapha.


Palloptera is a genus of flutter flies in the family Pallopteridae. There are at least 30 described species in Palloptera.


Pallopteridae is a family of flies. The various species are collectively called flutter-wing flies, trembling-wing, or waving-wing flies, because of the striking vibration of the wings in many species. Over 70 species in about 15 genera are found in the temperate regions of the Northern and Southern Hemispheres.


Piophila is a genus of small two-winged flies which includes the species known as the cheese fly. As a whole, the genus is often called wine flies; they prefer fermented foodstuffs to deposit their eggs. Piophila vulgaris visits Arum maculatum flowers in England.

The genus is found in the Palearctic.


Platystoma is a genus of flies (Diptera) belonging to the family Platystomatidae.


The Platystomatidae (signal flies) are a distinctive family of flies (Diptera) in the superfamily Tephritoidea.

Signal flies are worldwide in distribution, found in all the ecozones, but predominate in the tropics. It is one of the larger families of acalyptrate Diptera with around 1200 species in 119 genera.


Prochyliza is a genus of waltzing flies in the family Piophilidae. There are about 10 described species in Prochyliza.


The Pyrgotidae are an unusual family of flies (Diptera), one of only two families of Cyclorrhapha that lack ocelli. Most species are "picture-winged" (i.e, have patterns of bands or spots on the wings), as is typical among the Tephritoidea, but unlike other tephritoids, they are endoparasitoids; the females pursue scarab beetles in flight, laying an egg on the beetle's back under the elytra where the beetle cannot reach it. The egg hatches and the fly larva enters the body cavity of the beetle, feeding and eventually killing the host before pupating. In the United States, some species of Pyrgota and Sphecomyiella can be quite common in areas where their host beetles (typically the genus Phyllophaga, or "June beetles") are abundant. Like their host beetles, these flies are primarily nocturnal, and are often attracted to artificial lights.


The Richardiidae are a family of Diptera in the superfamily Tephritoidea.

This small family consists of just over 30 genera and 175 species. Almost all species are neotropical. Generally, the biology of the richardiids is little known. Some of the larvae are plant feeders or saprophages in decaying plant material. One species, the pineapple fruit fly Melanoloma viatrix, has been reported as a pest of pineapples. Most adults have some general features, conspicuously pictured wings, often with metallic blue or greenish colors on the body and legs, and a typical tephritoid ovipositor.


Rivellia is a genus of signal flies (insects in the family Platystomatidae). There are at least 140 described species in Rivellia.


Senopterina is a genus of signal flies (insects in the family Platystomatidae). There are about 17 described species in Senopterina.


The Ulidiidae (formerly Otitidae) or picture-winged flies are a large and diverse cosmopolitan family of flies (Diptera), and as in related families, most species are herbivorous or detritivore. They are often known as picture-winged flies, along with members of other families in the superfamily Tephritoidea that have patterns of bands or spots on the wings. Some species share with the Tephritidae an unusual elongated posteroapical projection of the anal cell in the wing, but can be differentiated by the smoothly curving subcostal vein. Two species, Tetanops myopaeformis and Euxesta stigmatias, are agricultural pests.

Extant Diptera families


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