In biology, a taxon (plural taxa; back-formation from taxonomy) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and given a particular ranking, especially if and when it is accepted or becomes established. It is not uncommon, however, for taxonomists to remain at odds over what belongs to a taxon and the criteria used for inclusion. If a taxon is given a formal scientific name, its use is then governed by one of the nomenclature codes specifying which scientific name is correct for a particular grouping.
Initial attempts at classifying and ordering organisms (plants and/or animals) was set forth in Linnaeus's system in Systema Naturae, 10th edition, as well as an unpublished work by Bernard and Antoine Laurent de Jussieu. The idea of a unit-based system of biological classification was first made widely available in 1805 in the introduction of Jean-Baptiste Lamarck's Flore françoise, of Augustin Pyramus de Candolle's Principes élémentaires de botanique. Lamarck set out a system for the "natural classification" of plants. Since then, systematists continue to construct accurate classifications encompassing the diversity of life; today, a "good" or "useful" taxon is commonly taken to be one that reflects evolutionary relationships.[n 1]
Many modern systematists, such as advocates of phylogenetic nomenclature, use cladistic methods that require taxa to be monophyletic (all descendants of some ancestor). Their basic unit, therefore, is the clade rather than the taxon. Similarly, among those contemporary taxonomists working with the traditional Linnean (binomial) nomenclature, few propose taxa they know to be paraphyletic. An example of a well-established taxon that is not also a clade is the class Reptilia, the reptiles; birds are descendants of reptiles but are not included in the Reptilia.
The term taxon was first used in 1926 by Adolf Meyer-Abich for animal groups, as a backformation from the word Taxonomy; the word Taxonomy had been coined a century before from the Greek components τάξις (taxis, meaning arrangement) and -νομία (-nomia meaning method). For plants, it was proposed by Herman Johannes Lam in 1948, and it was adopted at the VII International Botanical Congress, held in 1950.
A taxon can be assigned a taxonomic rank, usually (but not necessarily) when it is given a formal name.
A prefix is used to indicate a ranking of lesser importance. The prefix super- indicates a rank above, the prefix sub- indicates a rank below. In zoology the prefix infra- indicates a rank below sub-. For instance, among the additional ranks of class are superclass, subclass and infraclass.
Rank is relative, and restricted to a particular systematic schema. For example, liverworts have been grouped, in various systems of classification, as a family, order, class, or division (phylum). The use of a narrow set of ranks is challenged by users of cladistics; for example, the mere 10 ranks traditionally used between animal families (governed by the ICZN) and animal phyla (usually the highest relevant rank in taxonomic work) often cannot adequately represent the evolutionary history as more about a lineage's phylogeny becomes known.
In addition, the class rank is quite often not an evolutionary but a phenetic or paraphyletic group and as opposed to those ranks governed by the ICZN (family-level, genus-level and species-level taxa), can usually not be made monophyletic by exchanging the taxa contained therein. This has given rise to phylogenetic taxonomy and the ongoing development of the PhyloCode, which has been proposed as a new alternative to replace Linnean classification and govern the application of names to clades. Many cladists do not see any need to depart from traditional nomenclature as governed by the ICZN, ICN, etc.
La taxinomie s'enrichit avec l'invenition du mot «taxon» par Adolf Meyer-Abich, naturaliste allemand, dans sa Logik der morphologie, im Rahmen einer Logik der gesamten Biologie (1926) [Translation: Taxonomy is enriched by the invention of the word "taxon" by Adolf Meyer-Abich, German naturalist, in his Logik der morphologie, im Rahmen einer Logik der gesamten Biologie (1926).]CS1 maint: Multiple names: authors list (link)
Acacia, commonly known as the wattles or acacias, is a large genus of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially it comprised a group of plant species native to Africa and Australia, with the first species A. nilotica described by Linnaeus. Controversy erupted in the early 2000s when it became evident that the genus as it stood was not monophyletic, and that several divergent lineages needed to be placed in separate genera. It turned out that one lineage comprising over 900 species mainly native to Australia was not closely related to the mainly African lineage that contained A. nilotica—the first and type species. This meant that the Australian lineage (by far the most prolific in number of species) would need to be renamed. Botanist Les Pedley named this group Racosperma, which was inconsistently adopted. Australian botanists proposed that this would be more disruptive than setting a different type species (A. penninervis) and allowing this large number of species to remain Acacia, resulting in the two African lineages being renamed Vachellia and Senegalia, and the two New World lineages renamed Acaciella and Mariosousa. This was officially adopted, but many botanists from Africa and elsewhere disagreed that this was necessary.
A number of species have been introduced to various parts of the world, and two million hectares of commercial plantations have been established. The heterogeneous group varies considerably in habit, from mat-like subshrubs to canopy trees in forest.Crayfish
Crayfish, also known as crawfish, crawdads, crawlfish, crawldads, freshwater lobsters, mountain lobsters, mudbugs, or yabbies are freshwater crustaceans resembling small lobsters (to which they are related). Taxonomically, they are members of the superfamilies Astacoidea and Parastacoidea. They breathe through feather-like gills. Some species are found in brooks and streams where there is running fresh water, while others thrive in swamps, ditches, and paddy fields. Most crayfish cannot tolerate polluted water, although some species such as Procambarus clarkii are hardier. Crayfish feed on animals and plants, either living or decomposing, and detritus.Extinction
In biology, extinction is the termination of an organism or of a group of organisms (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the fossil record) after a period of apparent absence.
More than 99 percent of all species, amounting to over five billion species, that ever lived on Earth are estimated to have died out. Estimates on the number of Earth's current species range from 10 million to 14 million, of which about 1.2 million have been documented and over 86 percent have not yet been described. In 2016, scientists reported that 1 trillion species are estimated to be on Earth currently with only one-thousandth of one percent described.Through evolution, species arise through the process of speciation—where new varieties of organisms arise and thrive when they are able to find and exploit an ecological niche—and species become extinct when they are no longer able to survive in changing conditions or against superior competition. The relationship between animals and their ecological niches has been firmly established. A typical species becomes extinct within 10 million years of its first appearance, although some species, called living fossils, survive with virtually no morphological change for hundreds of millions of years.
Mass extinctions are relatively rare events; however, isolated extinctions are quite common. Only recently have extinctions been recorded and scientists have become alarmed at the current high rate of extinctions. Most species that become extinct are never scientifically documented. Some scientists estimate that up to half of presently existing plant and animal species may become extinct by 2100. A 2018 report indicated that the phylogenetic diversity of 300 mammalian species erased during the human era since the Late Pleistocene would require 5 to 7 million years to recover.A dagger symbol (†) placed next to the name of a species or other taxon normally indicates its status as extinct.Genus
A genus (, pl. genera ) is a taxonomic rank used in the biological classification of living and fossil organisms, as well as viruses, in biology. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus.
E.g. Panthera leo (lion) and Panthera onca (jaguar) are two species within the genus Panthera. Panthera is a genus within the family Felidae.The composition of a genus is determined by a taxonomist. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful:
monophyly – all descendants of an ancestral taxon are grouped together (i.e. phylogenetic analysis should clearly demonstrate both monophyly and validity as a separate lineage).
reasonable compactness – a genus should not be expanded needlessly; and
distinctness – with respect to evolutionarily relevant criteria, i.e. ecology, morphology, or biogeography; DNA sequences are a consequence rather than a condition of diverging evolutionary lineages except in cases where they directly inhibit gene flow (e.g. postzygotic barriers).Moreover, genera should be composed of phylogenetic units of the same kind as other (analogous) genera.Incertae sedis
Incertae sedis (Latin for "of uncertain placement") or problematica are terms used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty at specific taxonomic levels is indicated by incertae familiae (of uncertain family), incerti subordinis (of uncertain suborder), incerti ordinis (of uncertain order) and similar terms.Interim Register of Marine and Nonmarine Genera
The Interim Register of Marine and Nonmarine Genera (IRMNG) is a taxonomic database containing the scientific names of the genus, species, and higher ranks of many plants, animals and other kingdoms, both living and extinct, within a standardized taxonomic hierarchy, with associated machine-readable information on habitat (e.g. marine/nonmarine) and extant/fossil status for the majority of entries. The database aspires to provide complete coverage of both accepted and unaccepted genus names across all kingdoms, with a subset only of species names included as a lower priority. In its March 2019 release, IRMNG contained 490,095 genus names, of which 236,514 were listed as "accepted", 120,194 "unaccepted", 7,391 of "other" status i.e. interim unpublished, nomen dubium, nomen nudum, taxon inquirendum or temporary name, and 125,996 as "uncertain" (unassessed for taxonomic status at this time). The data originate from a range of (frequently domain-specific) print, online and database sources, and are reorganised into a common data structure to support a variety of online queries, generation of individual taxon pages, and bulk data supply to other biodiversity informatics projects. IRMNG content can be queried and displayed freely via the web, and download files of the data down to the taxonomic rank of genus as at specific dates are available in the Darwin Core Archive (DwC-A) format.
The data include homonyms (with their authorities), including both available (validly published) and selected unavailable names.IRMNG was initiated in 2006 by the Ocean Biogeographic Information System (OBIS) Australia at CSIRO Marine and Atmospheric Research, and since has been hosted by the Flanders Marine Institute (VLIZ) since 2016. VLIZ also hosts the World Register of Marine Species (WoRMS), using a common infrastructure.Content from IRMNG is used by several global Biodiversity Informatics projects including Open Tree of Life, the Global Biodiversity Information Facility (GBIF), and the Encyclopedia of Life (EOL), in addition to others including the Atlas of Living Australia and the Global Names Architecture (GNA)'s Global Names Resolver. From 2018 onwards, IRMNG data are also being used to populate the taxonomic hierarchy and provide generic names for a range of taxa in the areas of protists (kingdoms Protozoa and Chromista) and plant algae (Charophyta, Chlorophyta, Glaucophyta and Rhodophyta) in the Catalogue of Life. IRMNG identifiers have also been associated with numerous Wikipedia taxon pages, based on content harvested from IRMNG and stored in Wikidata.IRMNG was initiated and designed by Tony Rees. For his work on this and other projects, GBIF awarded him the 2014 Ebbe Nielsen Prize. The citation said, in part:
IRMNG in particular has been a tool of enormous importance to GBIF and others in supplying much of the detail for a global taxonomic classification of all life and as high-value taxon trait data in a form which can readily be reused in data validation and to enhance species pages.
IRMNG is now managed and curated by Rees, with assistance from the VLIZ team.Lemon
The lemon, Citrus limon (L.) Osbeck, is a species of small evergreen tree in the flowering plant family Rutaceae, native to South Asia, primarily North eastern India.
The tree's ellipsoidal yellow fruit is used for culinary and non-culinary purposes throughout the world, primarily for its juice, which has both culinary and cleaning uses. The pulp and rind (zest) are also used in cooking and baking. The juice of the lemon is about 5% to 6% citric acid, with a pH of around 2.2, giving it a sour taste. The distinctive sour taste of lemon juice makes it a key ingredient in drinks and foods such as lemonade and lemon meringue pie.Monophyly
In cladistics, a monophyletic group, or clade, is a group of organisms that consists of all the descendants of a common ancestor. Monophyletic groups are typically characterised by shared derived characteristics (synapomorphies), which distinguish organisms in the clade from other organisms. The arrangement of the members of a monophyletic group is called a monophyly.
Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A paraphyletic group consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A polyphyletic group is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor.
These definitions have taken some time to be accepted. When the cladistics school of thought became mainstream in the 1960s, several alternative definitions were in use. Indeed, taxonomists sometimes used terms without defining them, leading to confusion in the early literature, a confusion which persists.The first diagram shows a phylogenetic tree with two monophyletic groups. The several groups and subgroups are particularly situated as branches of the tree to indicate ordered lineal relationships between all the organisms shown. Further, any group may (or may not) be considered a taxon by modern systematics, depending upon the selection of its members in relation to their common ancestor(s); see second and third diagrams.Monotypic taxon
In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon.A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispecific" or "monospecific" is sometimes preferred.
In botanical nomenclature, a monotypic genus is a genus in the special case where a genus and a single species are simultaneously described.In contrast an oligotypic taxon contains more than one but only a very few subordinate taxa.Neontology
Neontology is a part of biology that, in contrast to paleontology, deals with living (or, more generally, recent) organisms. It is the study of extant taxa (singular: extant taxon): taxa (such as species, genera and families) with members still alive, as opposed to (all) being extinct. For example:
The moose (Alces alces) is an extant species, and the dodo is an extinct species.
In the group of molluscs known as the cephalopods, as of 1987 there were approximately 600 extant species and 7,500 extinct species.A taxon can be classified as extinct if it is broadly agreed or certified that no members of the group are still alive. Conversely, an extinct taxon can be reclassified as extant if there are new discoveries of extant species ("Lazarus species"), or if previously-known extant species are reclassified as members of the taxon.
Most biologists, zoologists, and botanists are in practice neontologists, and the term neontologist is used largely by paleontologists referring to non-paleontologists. Stephen Jay Gould said of neontology:
All professions maintain their parochialisms, and I trust that nonpaleontological readers will forgive our major manifestation. We are paleontologists, so we need a name to contrast ourselves with all you folks who study modern organisms in human or ecological time. You therefore become neontologists. We do recognize the unbalanced and parochial nature of this dichotomous division.
Neontological evolutionary biology has a temporal perspective between 100 to 1000 years. Neontology's fundamental basis relies on models of natural selection as well as speciation. Neontology's methods, when compared to evolutionary paleontology, has a greater emphasis on experiments. There are more frequent discontinuities present in paleontology than in neontology, because paleontology involves extinct taxa. Neontology has organisms actually present and available to sample and perform research on. Neontology's research method uses cladistics to examine morphologies and genetics. Neontology data has more emphasis on genetic data and the population structure than paleontology does.Pine
A pine is any conifer in the genus Pinus () of the family Pinaceae. Pinus is the sole genus in the subfamily Pinoideae. The Plant List compiled by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accepts 126 species names of pines as current, together with 35 unresolved species and many more synonyms.Rose
A rose is a woody perennial flowering plant of the genus Rosa, in the family Rosaceae, or the flower it bears. There are over three hundred species and thousands of cultivars. They form a group of plants that can be erect shrubs, climbing, or trailing, with stems that are often armed with sharp prickles. Flowers vary in size and shape and are usually large and showy, in colours ranging from white through yellows and reds. Most species are native to Asia, with smaller numbers native to Europe, North America, and northwestern Africa. Species, cultivars and hybrids are all widely grown for their beauty and often are fragrant. Roses have acquired cultural significance in many societies. Rose plants range in size from compact, miniature roses, to climbers that can reach seven meters in height. Different species hybridize easily, and this has been used in the development of the wide range of garden roses.The name rose comes from French, itself from Latin rosa, which was perhaps borrowed from Oscan, from Greek ρόδον rhódon (Aeolic βρόδον wródon), itself borrowed from Old Persian wrd- (wurdi), related to Avestan varəδa, Sogdian ward, Parthian wâr.Sister group
A sister group or sister taxon is a phylogenetic term denoting the closest relatives of another given unit in an evolutionary tree. The expression is most easily illustrated by a cladogram:
The sister group to A is B; conversely, the sister group to B is A. Groups A and B, together with all other descendants of their most recent common ancestor, form the clade AB. The sister group to clade AB is C.
The whole clade ABC is itself a subtree of a larger tree, which offers yet more sister group branches that are related but farther removed from the leaf nodes, such as A, B, and C.
In cladistic standards, A, B, and C may represent specimens, species, taxon-groups, etc. If they represent species, the term sister species is sometimes used.
The term "sister group" is used in phylogenetic analysis, and only groups identified in the analysis are labeled as sister groups. An example is in birds, whose sister group is commonly cited as the crocodiles, but that is true only when dealing with extant taxa. The bird family tree is rooted in the dinosaurs, making for a number of extinct groups branching off before coming to the last common ancestor of birds and crocodiles. Thus, the term sister group must be seen as a relative term, with the caveat that the sister group is the closest relative only among the groups/species/specimens that are included in the analysis.Species description
A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species which have been described previously or are related. The species description often contains photographs or other illustrations of the type material and states in which museums it has been deposited. The publication in which the species is described gives the new species a formal scientific name. Some 1.9 million species have been identified and described, out of some 8.7 million that may actually exist. Millions more have become extinct.Synonym (taxonomy)
In scientific nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name, although the term is used somewhat differently in the zoological code of nomenclature. For example, Linnaeus was the first to give a scientific name (under the currently used system of scientific nomenclature) to the Norway spruce, which he called Pinus abies. This name is no longer in use: it is now a synonym of the current scientific name, Picea abies.
Unlike synonyms in other contexts, in taxonomy a synonym is not interchangeable with the name of which it is a synonym. In taxonomy, synonyms are not equals, but have a different status. For any taxon with a particular circumscription, position, and rank, only one scientific name is considered to be the correct one at any given time (this correct name is to be determined by applying the relevant code of nomenclature). A synonym cannot exist in isolation: it is always an alternative to a different scientific name. Given that the correct name of a taxon depends on the taxonomic viewpoint used (resulting in a particular circumscription, position and rank) a name that is one taxonomist's synonym may be another taxonomist's correct name (and vice versa).
Synonyms may arise whenever the same taxon is described and named more than once, independently. They may also arise when existing taxa are changed, as when two taxa are joined to become one, a species is moved to a different genus, a variety is moved to a different species, etc. Synonyms also come about when the codes of nomenclature change, so that older names are no longer acceptable; for example, Erica herbacea L. has been rejected in favour of Erica carnea L. and is thus its synonym.Taxonomic rank
In biological classification, taxonomic rank is the relative level of a group of organisms (a taxon) in a taxonomic hierarchy. Examples of taxonomic ranks are species, genus, family, order, class, phylum, kingdom, domain, etc.
A given rank subsumes under it less general categories, that is, more specific descriptions of life forms. Above it, each rank is classified within more general categories of organisms and groups of organisms related to each other through inheritance of traits or features from common ancestors. The rank of any species and the description of its genus is basic; which means that to identify a particular organism, it is usually not necessary to specify ranks other than these first two.Consider a particular species, the red fox, Vulpes vulpes: the next rank above, the genus Vulpes, comprises all the "true" foxes. Their closest relatives are in the immediately higher rank, the family Canidae, which includes dogs, wolves, jackals, and all foxes; the next higher rank, the order Carnivora, includes caniforms (bears, seals, weasels, skunks, raccoons and all those mentioned above), and feliforms (cats, civets, hyenas, mongooses). Carnivorans are one group of the hairy, warm-blooded, nursing members of the class Mammalia, which are classified among animals with backbones in the phylum Chordata, and with them among all animals in the kingdom Animalia. Finally, at the highest rank all of these are grouped together with all other organisms possessing cell nuclei in the domain Eukarya.
The International Code of Zoological Nomenclature defines rank as: "The level, for nomenclatural purposes, of a taxon in a taxonomic hierarchy (e.g. all families are for nomenclatural purposes at the same rank, which lies between superfamily and subfamily)."Taxonomy (biology)
In biology, taxonomy (from Ancient Greek τάξις (taxis), meaning 'arrangement', and -νομία (-nomia), meaning 'method') is the science of defining and naming groups of biological organisms on the basis of shared characteristics. Organisms are grouped together into taxa (singular: taxon) and these groups are given a taxonomic rank; groups of a given rank can be aggregated to form a super-group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are domain, kingdom, phylum (division is sometimes used in botany in place of phylum), class, order, family, genus, and species. The Swedish botanist Carl Linnaeus is regarded as the father of taxonomy, as he developed a system known as Linnaean taxonomy for categorizing organisms and binomial nomenclature for naming organisms.
With the advent of such fields of study as phylogenetics, cladistics, and systematics, the Linnaean system has progressed to a system of modern biological classification based on the evolutionary relationships between organisms, both living and extinct.Type (biology)
In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralize the defining features of that particular taxon. In older usage (pre-1900 in botany), a type was a taxon rather than a specimen.A taxon is a scientifically named grouping of organisms with other like organisms, a set that includes some organisms and excludes others, based on a detailed published description (for example a species description) and on the provision of type material, which is usually available to scientists for examination in a major museum research collection, or similar institution.Watermelon
Citrullus lanatus is a plant species in the family Cucurbitaceae, a vine-like (scrambler and trailer) flowering plant originating in West Africa. It is cultivated for its fruit. The subdivision of this species into two varieties, watermelons (Citrullus lanatus (Thunb.) var. lanatus) and citron melons (Citrullus lanatus var. citroides (L. H. Bailey) Mansf.), originated with the erroneous synonymization of Citrullus lanatus (Thunb.) Matsum. & Nakai and Citrullus vulgaris Schrad. by L.H. Bailey in 1930. Molecular data including sequences from the original collection of Thunberg and other relevant type material, show that the sweet watermelon (Citrullus vulgaris Schrad.) and the bitter wooly melon Citrullus lanatus (Thunb.) Matsum. & Nakai are not closely related to each other. Since 1930, thousands of papers have misapplied the name Citrullus lanatus (Thunb.) Matsum. & Nakai for the watermelon, and a proposal to conserve the name with this meaning was accepted by the relevant nomenclatural committee and confirmed at the International Botanical Congress in Shenzhen in China in 2017.The bitter South African melon first collected by Thunberg has become naturalized in semiarid regions of several continents, and is designated as a "pest plant" in parts of Western Australia where they are called pig melon.Watermelon (Citrullus lanatus) is a scrambling and trailing vine in the flowering plant family Cucurbitaceae. The species was long thought to have originated in southern Africa, but this was based on the erroneous synonymization by L. H. Bailey (1930) of a South African species with the cultivated watermelon. The error became apparent with DNA comparison of material of the cultivated watermelon seen and named by Linnaeus and the holotype of the South African species. There is evidence from seeds in Pharaoh tombs of watermelon cultivation in Ancient Egypt. Watermelon is grown in tropical and subtropical areas worldwide for its large edible fruit, also known as a watermelon, which is a special kind of berry with a hard rind and no internal division, botanically called a pepo. The sweet, juicy flesh is usually deep red to pink, with many black seeds, although seedless varieties have been cultivated. The fruit can be eaten raw or pickled and the rind is edible after cooking.
Considerable breeding effort has been put into disease-resistant varieties. Many cultivars are available that produce mature fruit within 100 days of planting the crop.
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