Stomatosuchus inermis is an extinct 10 metres (33 ft) long stomatosuchid neosuchian from the Late Cretaceous (Cenomanian) of Egypt. Unlike most other crocodyliforms, it is difficult to determine exactly what S. inermis ate. Its flattened skull had a long, flat, lid-like snout, which was lined with small, conical teeth. The mandible may have been toothless and may have supported a pelican-like throat pouch.[1]

The only known specimen, a large skull, which was collected in German paleontologist Ernst Stromer's Egyptian expedition, was obliterated when the Munich Museum was destroyed during an Allied bombing raid in 1944.[2][3]

Temporal range: 100–94 Ma
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Family: Stomatosuchidae
Genus: Stomatosuchus
Stromer, 1925
Type species
Stomatosuchus inermis
Stromer, 1925
Stomatosuchus inermis
Comparison of Stomatosuchus (center) with Laganosuchus (bottom)


  1. ^ Naish, D. 2002. Fossils explained 34: Crocodilians. Geology Today 2: 71-77.
  2. ^ Stromer, E. (1925). Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wüsten Ägyptens. II. Wirbeltier-Reste der Baharije-Stufe (unterstes Cenoman). 7. Stomatosuchus inermis Stromer, ein schwach bezahnter Krokodilier und 8. Ein Skelettrest des Pristiden Onchopristis numidus Haug sp. Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Abteilung 30(6): 1–22.
  3. ^ Sereno, P. C.; Larsson, H. C. E. (2009). "Cretaceous crocodyliforms from the Sahara". ZooKeys. 28 (2009): 1–143. doi:10.3897/zookeys.28.325.

Aegyptosuchidae is an extinct family of eusuchian crocodyliforms from the Cretaceous period of Africa. They are characterized by their large size and flat heads. The family includes two genera, Aegyptosuchus and Aegisuchus.


Atoposauridae is a family of crocodile-line archosaurs belonging to Neosuchia. The majority of the family are known from Late Jurassic to Early Cretaceous marine deposits in France, Portugal, and Bavaria in southern Germany. The discovery of the genus Aprosuchus, however, extends the duration of the lineage to the end of the Cretaceous in Romania.

Bahariya Formation

The Bahariya Formation (also transcribed as Baharija Formation) is a fossiliferous geologic formation dating back to the Early Cenomanian, which outcrops within the Bahariya depression in Egypt, and is known from oil exploration drilling across much of the Western Desert where it forms an important oil reservoir.


Brillanceausuchus is an extinct genus of atoposaurid crocodylomorph. Fossils have been found in Early Cretaceous–age rocks of Cameroon. The genus is notable for the position of the secondary choana within its palate. Parts of the pterygoid bones make up the rostral margin of the choana and thus separate it from the palatines, a feature also seen in the more advanced neosuchian suborder Eusuchia. This characteristic was once thought to be characteristic of Eusuchia, but its presence in Brillanceausuchus suggests that the trait is homoplasic, thus making the evolution of the position of the choana within crocodilians more complex than previously thought.


Chiayusuchus is an extinct genus of stomatosuchid crocodylomorph. Fossils have been found from the Xinminbao Group in Gansu, China. It existed during the Barremian and Aptian stages of the Early Cretaceous. It was in the same family as the large, flat-snouted crocodilian Stomatosuchus.


Coelosuchus is an extinct genus of goniopholidid mesoeucrocodylian. Fossils have been found from the Graneros Shale of the Benton Group in Wyoming, and are of Cenomanian age. It was slightly over 1 meter in length.

Ernst Stromer

Ernst Freiherr Stromer von Reichenbach (12 June 1871 – in Nürnberg; 18 December 1952 in Erlangen) was a German paleontologist.

He described the following Cretaceous dinosaurs from Egypt: Aegyptosaurus, Bahariasaurus, Carcharodontosaurus, and the enigmatic theropod, Spinosaurus aegyptiacus. Stromer also described the giant crocodilian Stomatosuchus. The fossil bird genus Stromeria, named in his honor by Kálmán Lambrecht in 1929, is today synonymized with Eremopezus. The sauropod Paralititan stromeri is also named in his honor. The majority of his fossil discoveries were destroyed during World War Two and only photographs of some remain.


The Eusuchia ("true crocodiles") are a clade of crocodylomorphs that first appears in the Early Cretaceous with Hylaeochampsa. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survived the K-T extinction. Since the other two clades died out 35 and 11 million years ago, all living crocodilian species are eusuchians, as are many extinct forms.


Karatausuchus is an extinct genus of atoposaurid crocodylomorph. It is known from a single specimen discovered in the Late Jurassic (Oxfordian - Kimmeridgian) Karabastau Svita from the vicinity of Mikhailovka in the Karatau Mountains of southern Kazakhstan. The type specimen is PIN 25858/1, a complete but poorly preserved juvenile skeleton with some possible soft tissue preservation. It is notable for having over 90 teeth, but its other anatomical details are difficult to discern. The length of this individual is estimated at 160 millimetres (6.3 in). Karatausuchus was described in 1976 by Mikhail Efimov, and the type species is K. sharovi.


Khoratosuchus is an extinct genus of neosuchian crocodylomorph which existed in northeast Thailand during the early Cretaceous period. Its type species is Khoratosuchus jintasakuli. Khoratosuchus is the youngest and most advanced Mesozoic crocodyliform yet known from Thailand. It possesses several distinctive features that help determine its phylogenetic position among crocodylomorphs, including secondary choanae relatively posterior and almost encircled by the pterygoid bones on the palate and a smooth dorsal surface of the skull.


Libycosuchus is an extinct genus of North African crocodylomorph possibly related to Notosuchus. It was terrestrial, living approximately 95 million years ago in the Cenomanian stage of the Late Cretaceous. Fossil remains have been found in the Bahariya Formation in Egypt, making it contemporaneous with the crocodilian Stomatosuchus, and dinosaurs, including Spinosaurus. It was one of the few fossils discovered by Ernst Stromer that wasn't destroyed by the Royal Air Force during the bombing of Munich in 1944.


Mourasuchus is an extinct genus of giant, aberrant caiman from the Miocene of South America. Its skull has been described as duck like, being broad, flat and very elongate, closely resembling what is seen in Stomatosuchus, an unrelated crocodylian that may also have had a large gular sac similar to those of pelicans or baleen whales. Mourasuchus, is a strange nettosuchid with an unusually long, broad snout.The synonym Carandaisuchus was used to describe fossils from the Ituzaingó Formation in Argentina demonstrating a pronounced bony occipital crest.


Nannosuchus (meaning "dwarf crocodile") is an extinct genus of goniopholidid mesoeucrocodylian from the Berriasian of England.


Sabresuchus is an extinct genus of neosuchian crocodyliform from the Cretaceous of Europe. The name is derived from 'Sabre' in reference to the enlarged and curved fifth maxillary tooth, and 'suchus' from the Ancient Greek for crocodile.


Stomatosuchidae is an extinct family of neosuchian crocodylomorphs. It is defined as the most inclusive clade containing Stomatosuchus inermis but not Notosuchus terrestris, Simosuchus clarki, Araripesuchus gomesii, Baurusuchus pachecoi, Peirosaurus torminni, or Crocodylus niloticus. Two genera are known to belong to Stomatosuchidae: Stomatosuchus, the type genus, and Laganosuchus. Fossils have been found from Egypt, Morocco, and Niger. Both lived during the Cenomanian stage of the Late Cretaceous. The skulls of stomatosuchids are said to be platyrostral because they have unusually flattened, elongate, duck-shaped craniums with U-shaped jaws. This platyrostral condition is similar to what is seen in the "nettosuchid" Mourasuchus, which is not closely related to stomatosuchids as it is a more derived alligatoroid that existed during the Miocene.

Unlike Mourasuchus, stomatosuchids have jaws that are less strongly bowed. Additionally, the glenoid is rounded rather than cupped at the posterior end of the jaw, and the retroarticular process is straight rather than dorsally curving like in Mourasuchus and other extant crocodylians.The only existing specimens of stomatosuchids belong to the recently described genus Laganosuchus, which is known from two species, L. thaumastos and L. maghrebensis from the Echkar Formation in Niger and the Kem Kem Beds in Morocco, respectively. The genus Stomatosuchus is known only from a holotype skull collected from the Bahariya Formation in Egypt, which was destroyed in World War II with the bombing of the Munich Museum. Because Stomatosuchus is known only from brief accounts by Ernst Stromer and Franz Nopcsa (1926) and no additional material has ever been found, the genus remains enigmatic.The genus Aegyptosuchus was once considered to be a member of Stomatosuchidae, but it is now placed within its own family, Aegyptosuchidae.


Stromerosuchus is a dubious genus of Late Cretaceous crocodyliform. Fragmentary remains have been found from the Cenomanian-age Bahariya Formation of Egypt. The genus was named in 1936 by Oskar Kuhn. It is named in honor of Ernst Stromer, the German paleontologist who found the fossils in the Bahariya Oasis in 1911 and described them in 1922. After their discovery, the fossils, along with many others found from Bahariya, were in the possession of the Egyptian Geological Survey. In 1922, the fossils were sent back to Stromer (who was in Germany at the time), but they were badly crushed in shipment from Egypt. Because the known remains are so poor, the genus is now regarded as a nomen dubium. Some material has been referred to the genera Aegyptosuchus and Stomatosuchus, both named by Stromer from the Bahariya material.


Symptosuchus is an extinct genus of goniopholidid mesoeucrocodylian. It is known from the Late Cretaceous of Argentina. Argentine paleontologist Florentino Ameghino named the genus in 1899, along with the type species S. contortidens. It was formally described by Carlos Rusconi in 1934.


Wahasuchus is a genus of extinct mesoeucrocodylian of the Middle Campanian age found in the Quseir Formation, Egypt. The generic name derives from the Arabic word واحة (waha), which means "oasis", and souchos from the Greek in honor of crocodile-headed god of ancient Egypt. The specific egyptensis (Lat.) means from Egypt.Fossils of skull and jaw fragments, dorsal vertebrae, and fragmentary appendicular remains have been recovered.


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