The nematoceran infraorder Psychodomorpha (sometimes misspelled Psychomorpha - which is also the name of a genus of noctuid moths) includes two common families, Psychodidae and Scatopsidae, and other very small, rare families. In some classifications (e.g. the Tree of Life Web Project), the group is paraphyletic.

Clogmia Albipunctata or moth fly
Clogmia albipunctata family Psychodidae
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Diptera
Suborder: Nematocera
Infraorder: Psychodomorpha
Hennig, 1968

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Alloeodidicrum is a genus of drain flies in the subfamily Psychodinae. It consists of two species.


Berdeniella is a genus of drain flies in the subfamily Psychodinae.


The Blephariceromorpha are an infraorder of nematoceran flies, including three families associated with fast-flowing, high-mountain streams, where the larvae can be found.

One recent classification based largely on fossils splits this group into two infraorders, and removes the Nymphomyiidae to its own suborder, but this has not gained wide acceptance. More recently, the family Blephariceridae has been considered a member of the infraorder Psychodomorpha, with Deuterophlebiidae and Nymphomyiidae either assigned their own infraorders or left unassigned to infraorder. The placement of these three families remains controversial.


The Canthyloscelidae are a small family of midges closely related to the Scatopsidae.

Adults are small to medium-sized (2.5-9.0 mm) flies, relatively stout, usually dark coloured Nematocera with stout legs. They are associated with ancient woodland. Larvae are xylosaprophagous and live in the moist, rotting wood of stumps and fallen trees.Most are considered endangered due to the vulnerability of their habitat.

There are 15 described species worldwide from New Zealand, North America, South America, Japan and Russia. The 3 species in Europe are suspected to be introductions. There is one know fossil species from the Jurassic.


Clogmia is a genus of drain flies in the subfamily Psychodinae.


Colobostema is a genus of minute black scavenger flies in the family Scatopsidae. There are at least 4 described species in Colobostema.


Ectaetia is a genus of minute black scavenger fly in the family Scatopsidae. Examples can be found in the Nearctic, Palearctic, Oriental, and Neotropical biogeographic realms. Adults of Ectaetia species are generally shiny, black, and large.


Exiliscelis is a genus of gnats, gall midges, and March flies in the family Canthyloscelidae. There is at least one described species in Exiliscelis, E. californiensis.


Nemopalpus is a genus of moth and sand flies in the family Psychodidae, with at least 30 described species. Nemopalpus is sometimes spelled "Nemapalpus".

Nemopalpus nearcticus

Nemopalpus nearcticus (sometimes Nemapalpus), the sugarfoot moth fly, is a species of nematoceran flies in the family Psychodidae. It is endemic in the United States.The IUCN conservation status of Nemopalpus nearcticus is "EN", endangered. The species faces a high risk of extinction in the near future.


Pericoma is a genus of moth flies in the family Psychodidae. There are at least 190 described species in Pericoma.


The Perissommatidae are a family of flies (Diptera) that was newly proposed in 1962. The family contains five species, four from Australia and one from Chile. The Perissommatidae are unusual as they have four eyes. They have a small slender body less than 2 mm in length. Their wings are large in comparison to their bodies and subsequently their flight is weak. Preferring high-altitude forest environments, adults only fly in the winter. In the case of Perissomma macalpinei, numbers of adults have been observed congregate in clumps of foliage, rising in short, zigzag flights in the sunlight above the foliage for short periods before descending.


Psychoda is a genus of moth flies in the family Psychodidae. There are at least 370 described species in Psychoda.

Psychoda alternata

Psychoda alternata is a species of moth flies in the family Psychodidae.


The Psychodinae are the nominate subfamily of moth flies (Psychodidae), also known as drain flies. Like most of their relatives, they are usually found in damp habitats; some occur in caves. The small larvae are aquatic or semiterrestrial; the imagines feed on plant sap and nectar or not at all. Psychodinae are found in almost every country on Earth, including some subantarctic islands.


The superfamily Scatopsoidea includes families: Canthyloscelidae, Perissommatidae and Scatopsidae.

Sycorax (fly)

Sycorax is a genus of moth flies and sand flies in the family Psychodidae. There are at least 40 described species in Sycorax.

Synneuron decipiens

Synneuron decipiens is a species of nematoceran flies in the family Canthyloscelidae.

Trichomyia lengleti

Trichomyia lengleti is an extinct species of Cretaceous flies found in the Charentes region of France.It was discovered in 2008 when x-ray technology was used to peer into opaque pieces of amber.

Extant Diptera families


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