The Platypezoidea are a superfamily of true flies of the section Aschiza. Their closest living relatives are the Syrphoidea, which, for example, contain the hoverflies. Like these, the adults do not burst open their pupal cases with a ptilinum when hatching, thus the Aschiza do not have the inverted-U-shaped suture above the antennae. They are, however, muscomorphs, thus have a particular type of pupal case resembling a rounded barrel and called puparium.

Phorid fly2
Pseudacteon sp. (Phoridae)
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Diptera
Section: Aschiza
Superfamily: Platypezoidea

5, see text




Five families are placed in the Platypezoidea, listed below in taxonomic sequence:

The Ironomyiidae, Lonchopteridae, and Phoridae are sometimes separated as Phoroidea. The reduced Platypezoidea thus created unites the two families of flat-footed flies.


  1. ^ McAlpine, D.K. (2008). "New extant species of ironic flies (Diptera: Ironomyiidae) with notes on ironomyiid morphology and relationships". Proceedings of the Linnean Society of New South Wales (Print). New South Wales: The Linnean Society of New South Wales. 129: 17–38.

Apocephalus is a genus of ant-decapitating flies (insects in the family Phoridae). There are at least 300 described species in Apocephalus.


The Aschiza are a section of the Brachycera. Two large families, the Syrphidae and the Phoridae, and a number of smaller taxa are in this group. They are similar to most of the familiar Muscomorpha with one notable exception; they do not possess a ptilinum, so lack the prominent ptilinal suture on the face as in other muscoid flies. They do still have a puparium with a circular emergence opening, but it is not as precisely ellipsoid in shape as is typical for other muscoids.

The term was first used by Eduard Becher .


Callomyia is a genus of flies in the family Platypezidae. Some species can be found in Belgium.


Calotarsa is a genus of flat-footed flies (insects in the family Platypezidae). There are about six described species in Calotarsa.


Diplonevra is a genus of scuttle flies (insects in the family Phoridae). There are at least eight described species in Diplonevra.


The Ironomyiidae, or ironic flies, are a small family of insects of the order Diptera. Historically, they have been included in the family Platypezidae, and includes three extant species and a number of extinct fossil species.


Lonchoptera is a genus of spear-winged flies (Lonchopteridae). Their common name refers to their subacute (pointed) wings, which have a distinct and sexually dimorphic venation.


The Lonchopteridae (spear-winged flies or pointed-wing flies) are a family of small (2–5 mm), slender, yellow to brownish-black Diptera, occurring all over the world. Their common name refers to their pointed wings, which have a distinct venation. Many are parthenogenic; males are very rare, however, at least in North American species, and have a somewhat different venation than do the females.

Spear-winged flies are common in moist, shady, grassy areas, where the larvae are found within decaying vegetation. One species, Lonchoptera bifurcata, is cosmopolitan in distribution, and may have been transported via shipments of vegetables.


Megaselia is a genus of flies in the family Phoridae.


The Metopininae are a subfamily of flies in the family Phoridae.


The Brachyceran infraorder Muscomorpha is a large and diverse group of flies, containing the bulk of the Brachycera, and, most of the known flies. It includes a number of the most familiar flies, such as the housefly, the fruit fly, and the blow fly. The antennae are short, usually three-segmented, with a dorsal arista. Their bodies are often highly setose, and the pattern of setae is often taxonomically important.

The larvae of muscomorphs (in the sense the name is used here; see below) have reduced head capsules, and the pupae are formed inside the exoskeleton of the last larval instar; exit from this puparium is by a circular line of weakness, and this pupal type is called "cyclorrhaphous"; this feature gives this group of flies their traditional name, Cyclorrhapha.


Nothomicrodon aztecarum is a species of Neotropical flies, originally described from a larva collected in 1924 from a carton nest of the ant Azteca trigona. It is the only species in the genus Nothomicrodon, but shows none of the features of a hoverfly larva, the family in which it was originally classified, and instead belongs in the family Phoridae.


The Opetiidae or flat-footed flies are members of a family of flies (insects of the Order Diptera). The family contains only five extant species in two genera, Opetia, and Puyehuemyia. the remaining taxa are known only from fossils.


Phalacrotophora is a genus of scuttle flies (insects in the family Phoridae). There are at least 50 described species in Phalacrotophora.


Phorinae is a subfamily of flies in the family Phoridae. There are at least 90 described species in Phorinae.


Platypezidae is a family of true flies of the superfamily Platypezoidea. The more than 250 species are found worldwide primarily in woodland habitats. A common name is flat-footed flies, but this is also used for the closely related Opetiidae which were included in the Platypezidae in former times.Some other genera formerly included here have been recognized as quite more distant and are nowadays placed in the asilomorph family Atelestidae.


Polyporivora is a genus of flat-footed flies (insects in the family Platypezidae). There are about eight described species in Polyporivora.


Pseudacteon is a genus of flies in the family Phoridae. There are over 70 described species of Pseudacteon fly. They are also known as ant-decapitating flies due to their parasitic larval stage. An egg is injected by the female fly into the shoulder joint of an ant worker. Soon after, the egg undergoes rapid inflation as it appears to absorb ant hemolymph. This first instar larva migrates into the ant head and consumes the jaw muscle and other tissues, leaving the mandibles hanging and preparing a future exit space. After about two weeks, Eventually, the ant worker is termed a "zombie" because the fly larva has effectively taken control. The worker leaves the nest and dies in the leaf litter or in a crack in the soil. As it dies, the ant's head falls off, apparently when the fly larva woreleases an enzyme that dissolves the membrane attaching the ant's head to its body. The fly pupates in the detached head capsule, requiring a further two weeks before emerging through the ant's mouth.


Puliciphora is a genus of scuttle flies (insects in the family Phoridae). There are at least 110 described species in Puliciphora.

Extant Diptera families


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