Plateosauria

Plateosauria is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to the Late Cretaceous. The name Plateosauria was first coined by Gustav Tornier in 1913.[1] The name afterwards fell out of use until the 1980s.

Plateosauria is a node-based taxon. In 1998, Paul Sereno defined Plateosauria as the last common ancestor of Plateosaurus engelhardti and Massospondylus carinatus, and its descendants.[2] Peter Galton and Paul Upchurch in 2004 used a different definition: the last common ancestor of Plateosaurus engelhardti and Jingshanosaurus xinwaensis, and its descendants. In their cladistic analysis the Plateosauria belonged to the Prosauropoda, and included the Plateosauridae subgroup. In Galton's and Upchurch's study also Coloradisaurus, Euskelosaurus, Jingshanosaurus, Massospondylus, Mussaurus, Sellosaurus, and Yunnanosaurus proved to be plateosaurians.[3]

However, recent cladistic analyses suggest that the Prosauropoda as traditionally defined is paraphyletic to sauropods.[4][5][6][7][8] Prosauropoda, as currently defined, is a synonym of Plateosauridae as both contain the same taxa by definition.

The following cladogram simplified after an analysis presented by Apaldetti and colleagues in 2011.[8]

 Plateosauria 

Plateosauridae

 Massopoda 

Riojasauridae

Ignavusaurus

Sarahsaurus

Massospondylidae

Yunnanosaurus

Jingshanosaurus

Seitaad

Anchisauria

The following cladogram simplified after an analysis presented by Blair McPhee and colleagues in 2014.[9]

Plateosauria

Ruehleia

Plateosauridae

 Massopoda 
 Riojasauridae 

Eucnemesaurus

Riojasaurus

Massospondylidae

 Sauropodiformes 

Yunnanosaurus

Jingshanosaurus

Seitaad

Anchisauria

Plateosauria
Temporal range: Late Triassic-Late Cretaceous, 225–66 Ma
Plateosaurus Skelett 2
Plateosaurus engelhardti skeleton
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Order: Saurischia
Suborder: Sauropodomorpha
Clade: Plateosauria
Tornier, 1913
Subgroups

References

  1. ^ Tornier, G., 1913, "Reptilia (Paläontologie)" In: Handwörterbuch Naturwissenschaften 8: 337-376
  2. ^ Sereno, P.C. (1998). "A rationale for phylogenetic definitions, with applications to the higher-level taxonomy of Dinosauria". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 210: 41–83.
  3. ^ Galton, P.M & Upchurch, P. (2004). "Prosauropoda". In D. B. Weishampel, P. Dodson, & H. Osmólska (eds.), The Dinosauria (second edition). University of California Press, Berkeley: 232–258.
  4. ^ Yates, Adam M. (2003). "Species taxonomy of the sauropodomorph dinosaurs from the Löwenstein Formation (Norian, Late Triassic) of Germany". Palaeontology. 46 (2): 317–337. doi:10.1111/j.0031-0239.2003.00301.x.
  5. ^ Yates, Adam M. (2007). "The first complete skull of the Triassic dinosaur Melanorosaurus Haughton (Sauropodomorpha: Anchisauria)". In Barrett & Batten (eds.), Evolution and Palaeobiology. 77: 9–55. ISBN 978-1-4051-6933-2.
  6. ^ Pol D.; Garrido A.; Cerda I.A. (2011). "A New Sauropodomorph Dinosaur from the Early Jurassic of Patagonia and the Origin and Evolution of the Sauropod-type Sacrum". PLoS ONE. 6 (1): e14572. Bibcode:2011PLoSO...614572P. doi:10.1371/journal.pone.0014572. PMC 3027623. PMID 21298087.
  7. ^ Fernando E. Novas; Martin D. Ezcurra; Sankar Chatterjee; T. S. Kutty (2011). "New dinosaur species from the Upper Triassic Upper Maleri and Lower Dharmaram formations of central India". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 101 (3–4): 333–349. doi:10.1017/S1755691011020093.
  8. ^ a b Cecilia Apaldetti, Ricardo N. Martinez, Oscar A. Alcober and Diego Pol (2011). "A New Basal Sauropodomorph (Dinosauria: Saurischia) from Quebrada del Barro Formation (Marayes-El Carrizal Basin), Northwestern Argentina". PLoS ONE. 6 (11): e26964. Bibcode:2011PLoSO...626964A. doi:10.1371/journal.pone.0026964. PMC 3212523. PMID 22096511.CS1 maint: Uses authors parameter (link)
  9. ^ McPhee, B. W.; Yates, A. M.; Choiniere, J. N.; Abdala, F. (2014). "The complete anatomy and phylogenetic relationships of Antetonitrus ingenipes(Sauropodiformes, Dinosauria): Implications for the origins of Sauropoda". Zoological Journal of the Linnean Society. 171: 151. doi:10.1111/zoj.12127.
Anchisauria

The Anchisauria were a clade of sauropodomorph dinosaurs that lived during the Late Triassic and Early Jurassic. The name Anchisauria was first used by Galton and Upchurch in the second edition of The Dinosauria. Galton and Upchurch assigned two families of dinosaurs to the Anchisauria: the Anchisauridae and the Melanorosauridae. The more common prosauropods Plateosaurus and Massospondylus were placed in the sister clade Plateosauria.

However, recent research indicates that Anchisaurus is closer to sauropods than traditional prosauropods; thus, Anchisauria would also include Sauropoda.The following cladogram simplified after an analysis presented by Blair McPhee and colleagues in 2014.

Arcusaurus

Arcusaurus is an extinct genus of sauropodomorph dinosaur from the Early Jurassic (Hettangian to Sinemurian stages) of South Africa.

Arcusaurus was first named by Adam Yates, Matthew Bonnan and Johann Neveling in 2011 and the type species is Arcusaurus pereirabdalorum. The generic name is derived from Latin arcus, "rainbow", a reference to the Rainbow Nation. The specific epithet honours Lucille Pereira and Fernando Abdala who discovered the fossils .

Arcusaurus is known from two fragmentary skeletons collected in March 2006 at the Spion Kop Heelbo site from the upper Elliot Formation in Senekal in Free State. The holotype, BP/1/6235, consists of a partial skull. Some limb bones and vertebrae are included in the material. Both specimens represented juvenile individuals. From detailed features the describers concluded these were not the young of either Aardonyx or Massospondylus.

A phylogenetic study of Arcusaurus found it to be a basal sauropodomorph, placing it as the sister taxon of Efraasia and all of the more derived sauropodomorphs. Since Efraasia is known from the Norian stage of the Late Triassic, the close relationship with Arcusaurus implies that there was a 35-million-year ghost lineage of sauropodomorphs stretching from Late Triassic forms to Arcusaurus. However, Arcusaurus possesses many features unique to more advanced groups included in the clade Plateosauria, raising doubts about the results of the phylogenetic analysis.

Averostra

Averostra, or "bird snouts", is a clade that includes most theropod dinosaurs that have a promaxillary fenestra (fenestra promaxillaris), an extra opening in the front outer side of the maxilla, the bone that makes up the upper jaw. Two groups of averostrans, the Ceratosauria and the Orionides, survived into the Cretaceous period. When the Cretaceous–Paleogene extinction event occurred, ceratosaurians and two groups of orionideans within the clade Coelurosauria, the Tyrannosauroidea and Maniraptoriformes, were still extant. Only one subgroup of maniraptoriformes, Aves, survived the extinction event and persisted to the present day.

Avetheropoda

Avetheropoda, or "bird theropods", is a clade that includes carnosaurians and coelurosaurs to the exclusion of other dinosaurs.

Cerapoda

Cerapoda ("ceratopsians and ornithopods") is a clade of the dinosaur order Ornithischia.

Gyposaurus

Gyposaurus (meaning "vulture lizard", referring to the outdated hypothesis that prosauropods were carnivores) is a genus of basal sauropodomorph dinosaur from the early Jurassic of South Africa. It is usually considered to represent juveniles of other prosauropods, but "G." sinensis is regarded as a possibly valid species in recent reviews of the prosauropods (Galton and Upchurch, 2004).

Ignavusaurus

Ignavusaurus is a genus of basal sauropodomorph dinosaur that lived during the Early Jurassic in what is now Lesotho. Its fossils were found in the Upper Elliot Formation which is probably Hettangian in age (around 200 million years ago). It was described on the basis of a partial, well preserved articulated skeleton. The type species, I. rachelis, was described in 2010 by Spanish palaeontologist F. Knoll.

Jingshanosaurus

Jingshanosaurus (meaning "Jingshan lizard") is a genus of sauropodomorph dinosaurs from the early Jurassic period.

Mussaurus

Mussaurus (meaning "mouse lizard") is a genus of herbivorous sauropodomorph dinosaur that lived in southern Argentina during the Late Triassic, about 215 million years ago. It receives its name from the small size of the skeletons of juvenile and infant individuals, which were once the only known specimens of the genus. However, since Mussaurus is now known from adult specimens, the name is something of a misnomer; adults possibly reached 6 metres (20 ft) in length and weighed more than 1,000 kilograms (2,200 lb). Mussaurus possesses anatomical features suggesting a close, possibly transitional evolutionary relationship with true sauropods.

Nambalia

Nambalia is a genus of basal sauropodomorph dinosaur. It lived during the Late Triassic period (late Norian to earliest Rhaetian) in what is now Andhra Pradesh, central India. It is known from the holotype ISI R273, parts 1-3, partially articulated postcranial material and from the paratypes ISI R273, parts 4-29, including partial postcrania of at least two individuals of different sizes found closely associated and one of them is nearly the same size as the

holotype. ISI R273 was discovered and collected from the Upper Maleri Formation within the Pranhita–Godavari Basin,

north of Nambal village. It was first named by Fernando E. Novas, Martin D. Ezcurra, Sankar Chatterjee and T. S. Kutty in 2011 and the type species is Nambalia roychowdhurii. The generic name is derived from the Indian village of Nambal which is close to the type locality. The specific name honors Dr. Roy Chowdhuri, for his research on the Triassic vertebrate faunas of India. A cladistic analysis by Novas et al. found that Nambalia is basal to Efraasia, Plateosauravus, Ruehleia and Plateosauria, but more derived than Thecodontosaurus, Pantydraco, and Guaibasauridae. Nambalia was found along with the plateosaurid Jaklapallisaurus, a guaibasaurid, and two basal dinosauriforms.

Orionides

Orionides is a clade of tetanuran theropod dinosaurs from the Middle Jurassic to the Present. The clade includes most theropod dinosaurs, including birds.

Plateosauridae

Plateosauridae is a family of plateosaurian sauropodomorphs from the Late Triassic of Europe. Although several dinosaurs have been classified as plateosaurids over the years, the family Plateosauridae is now restricted to Plateosaurus. In another study, Yates (2003) sunk Sellosaurus into Plateosaurus (as P. gracilis).

Pulanesaura

Pulanesaura is an extinct genus of basal sauropod known from the Early Jurassic (late Hettangian to Sinemurian) Upper Elliot Formation of the Free State, South Africa. It contains a single species, Pulanesaura eocollum, known from partial remains of at least two subadult to adult individuals.

Riojasauridae

Riojasauridae is a family of sauropod-like dinosaurs from the Upper Triassic. It is known primarily from the genera Riojasaurus and Eucnemesaurus. Sites containing Riojasauridae include the Lower Elliot Formation of Orange Free State, South Africa (where fossils of Eucnemesaurus have been found), and Ischigualasto, in La Rioja Province, Argentina ( where fossils of Riojasaurus have been recovered).

Sauropodomorpha

Sauropodomorpha ( SOR-ə-POD-ə-MOR-fə; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had long necks and tails, were quadrupedal, and became the largest animals to ever walk the Earth. The "prosauropods", which preceded the sauropods, were smaller and were often able to walk on two legs. The sauropodomorphs were the dominant terrestrial herbivores throughout much of the Mesozoic Era, from their origins in the mid-Triassic (approximately 230 Ma) until their decline and extinction at the end of the Cretaceous (approximately 66 Ma).

Seitaad

Seitaad is a genus of sauropodomorph dinosaur which lived during the lower Jurassic period in what is now southern Utah, United States.

Seitaad is known from an articulated partial postcranial holotype skeleton referred to as UMNH VP 18040. The skeleton is missing its head, neck and tail. It was collected from the Early Jurassic Navajo Sandstone, the uppermost unit of the Glen Canyon Group, dating to the Pliensbachian stage, near Comb Ridge, San Juan County. A phylogenetic study of Seitaad found it to be a plateosaur sauropodomorph, placing it in Massospondylidae or alternatively (a less probable position) in Plateosauridae, but its placement within the Plateosauria is not well understood. In a cladistic analysis, presented by Apaldetti and colleagues in November 2011, Seitaad was found to be within Massopoda, just outside Anchisauria.Seitaad was first described by Joseph J. W. Sertich and Mark A. Loewen in 2010 and the type species is Seitaad ruessi. The generic name is derived from Séít‘áád (Navajo language), a mythological sand monster from the Diné folklore who buried its victims in dunes. Seitaad appears to have been entombed by the collapse of a sand dune. The specific name honours Everett Ruess, a young artist, poet and naturalist, who mysteriously disappeared in 1934 while exploring southern Utah. Seitaad is the second basal sauropodomorph dinosaur to have been identified in North America.

Unaysauridae

Unaysauridae is a family of basal sauropodomorphs from the Late Triassic of India and Brazil.

Yimenosaurus

Yimenosaurus (meaning "Yiman reptile") is an extinct genus of plateosaurid sauropodomorph dinosaur that lived in China in the Early Jurassic. The genus was first named in 1990 by Ziqi Bai, Jie Yang and Guohui Wang, along with its type and only species, Yimenosaurus youngi. The species name honours renowned Chinese paleontologist Yang Zhongjian, the father of Chinese paleontology, known as C.C. Young in English. Known material includes the holotype, an almost complete skull and mandible, as well as incomplete cervical and dorsal vertebrae, a mostly complete sacrum, an ilium, ischia, partial ribs and complete femur, and a paratype, a well-preserved postcrania with a fragmentary skull.

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