Pamela C. Rasmussen

Pamela Cecile Rasmussen (born October 16, 1959) is a prominent American ornithologist and expert on Asian birds. She was formerly a research associate at the Smithsonian Institution in Washington, D.C., and is based at the Michigan State University. She is associated with other major centers of research in the United States and the United Kingdom.

Rasmussen's early research investigated South American seabirds and fossil birds from North America. She later specialised in Asian birds describing several new species and clarifying the status of others, particularly white-eyes and owls. More recently, she has been involved in large scale collaborations looking at patterns of global biodiversity, and has assessed the taxonomic status of South Asian vultures.

She was the main author of Birds of South Asia: The Ripley Guide, a landmark publication due to its greater geographical and species coverage compared to its predecessors. As a result of her study of museum bird specimens when researching for the book, she was instrumental in unveiling the extent of the theft from museums and fraudulent documentation perpetrated by eminent British ornithologist Richard Meinertzhagen.

Pamela C. Rasmussen
BornOctober 16, 1959 (age 59)
Alma materWalla Walla University
Scientific career
InstitutionsSmithsonian Institution
Michigan State University
InfluencesSidney Dillon Ripley

Early life and career

Rasmussen is the daughter of Helen Rasmussen, a Seventh-Day Adventist, whose husband, Chester Murray Rasmussen, a doctor, had left the family when Pamela and her sisters were young. Her interest in birds started when her mother bought her the junior edition of Oliver Austin's Birds of the World, and Pamela subsequently always chose to receive bird books as presents.[1]

She took her M.S. in 1983 at Walla Walla University, an Adventist-affiliated university in southeast Washington, and her Ph.D. at the University of Kansas in 1990, where she studied blue-eyed shags,[2][3] and was introduced to evolutionary theory, which had not been taught at her alma mater.[1][4]

Rasmussen is a visiting assistant professor of zoology, and assistant museum curator of mammalogy and ornithology, at Michigan State University, having formerly been a research associate for the eminent American ornithologist S. Dillon Ripley at the Smithsonian Institution in Washington, D.C. She is a member of the American Ornithological Society (AOS) Committee on Classification and Nomenclature, a scientific associate with the bird group of the British Natural History Museum zoology section at Tring, and an associate editor of The Ibis, the scientific journal of the British Ornithologists' Union.[4] Pamela Rasmussen is married to Dr Michael D. Gottfried, who is Curator of Paleontology, Associate Professor of Geology, and Director of the Center for Integrative Studies in General Science at MSU.[5]

Research highlights

South American seabirds

Rasmussen’s early work was largely focused on studies of the systematics, ecology and behaviour of Patagonian seabirds, notably cormorants. She studied plumage variations in juvenile blue-eyed, king and red-legged cormorants,[6][7] and used plumage and behavioural patterns to establish relationships between king and blue-eyed shags.[8][9][10] She also reviewed the fishing activity of olivaceous cormorants.[11]

Asian birds

Athene blewitti
An 1891 painting of the forest owlet, which was rediscovered by Rasmussen in 1997

Rasmussen described four new Asian bird species from her study of museum specimens. The Nicobar scops owl Otus alius,[12] the Sangihe scops owl Otus collari,[13] and the cinnabar hawk owl Ninox ios, a Sulawesi endemic,[14] all in 1998, and the Taiwan bush-warbler Bradypterus alishanensis in 2000.[15] She rediscovered the forest owlet Athene blewitti, which had not been seen since 1884, in western India,[16][17] previous searches by S. Dillon Ripley, Salim Ali and others having failed because they relied on fake documentation from Richard Meinertzhagen.[18][19] In November 1997, Rasmussen and Ben King of the American Museum of Natural History spent ten days unsuccessfully searching two east Indian locations before driving west to the site of another old specimen, where King spotted a small, chunky owl with short, heavily white-feathered legs and huge claws, which Rasmussen confirmed as the target species whilst the owl was videotaped and photographed.[19]

With her colleagues, she clarified the taxonomy of Indonesian white-eyes, establishing the specific status of the Sangihe white-eye Zosterops nehrkorni and the Seram white-eye Z. stalkeri[20] and confirmed the identity of the Serendib scops owl which had originally been discovered in Sri Lanka by local ornithologist Deepal Warakagoda.[21]

The Togian white-eye, identified as a new species by Rasmussen and her colleagues in 2008

The imperial pheasant is a rare bird found in the forests of Vietnam and Laos. Rasmussen and her co-workers used morphology, hybridisation experiments, and DNA analysis to show that this pheasant, previously thought critically endangered, is actually a naturally occurring hybrid between the Vietnamese pheasant Lophura hatinhensis and the subspecies annamensis of the silver pheasant L. nycthemera.[22]

A 2008 paper saw a return to white-eye taxonomy with the formal description of the Togian white-eye Zosterops somadikartai, an endemic species of the Togian Islands of Indonesia, which, unlike most of its relatives, lacks the white ring around the eye which give this group of birds its name.[23] Rasmussen noted that the Togian white-eye is distinctive not only in its appearance, but also in its lilting song, which sounds higher pitched and is less varied in frequency than the songs of its close relatives.[23]

Pamela Rasmussen’s interest in Asian birds led to her involvement in more specifically conservation-directed projects. Two Gyps vultures, the Indian white-rumped vulture, Gyps bengalensis, and the "long-billed vulture" suffered a 99 percent population decrease in South Asia due to poisoning by diclofenac, a veterinary drug that causes kidney failure in birds that have eaten the carcasses of treated cattle.[24][25] Rasmussen showed that there are two distinct species of long-billed vulture: the Indian vulture G. indicus and slender-billed vulture G. tenuirostris. This is important to conservation, since a captive-breeding program has been established to assist the recovery of at-risk vulture species.[26]


Long billed vulture
Indian vulture, a vulnerable species newly split as a result of Rasmussen's research into the genus Gyps

In 2005, Rasmussen was part of a large multi-institutional collaboration investigating biodiversity hotspots, which have a prominent role in conservation. The study assessed locations quantitatively for three criteria of bird diversity – species richness, the level of threat, and the number of endemic species. The results demonstrated that hotspots did not show the same geographical distribution for each factor. Only 2.5% of hotspot areas are common to all three aspects of diversity, with over 80% of hotspots registering on only one criterion. Each criterion explained less than 24% of the variation in the other factors, suggesting that even within a single taxonomic class, different mechanisms are responsible for the origin and maintenance of various aspects of diversity. Consequently, the different types of hotspots also vary greatly in their utility as conservation tools.[27]

Rasmussen's recent work has concentrated on further large-scale collaborations with the same group of institutions studying global patterns in biodiversity. A survey of species richness and geographical range size did not show the decrease in range size from temperate regions to the tropics that had been previously assumed;[28] although that pattern was largely true in the northern hemisphere, it did not appear to apply in the southern hemisphere.[29] Research evaluating the relationship between extinction and human impact showed that, after controlling for species richness, the best predictors of the global pattern of extinction risk are measures of human impact, with ecological factors being of secondary importance.[30] An examination of the distribution of rare and threatened vertebrate species, showed differing patterns for bird, mammal and amphibian species, which has consequences for hotspot-based conservation strategies.[31]

Other studies by Rasmussen and her international colleagues looked at the importance of energy availability,[32][33] and a 2007 paper showed that global patterns of spatial turnover are driven principally by widespread species rather than restricted ones. This complements other work, and helps to establish a unified model of how terrestrial biodiversity varies both within and between the Earth's major land masses.[34]


A fossil site at a borrow pit in near Cheswold, Delaware created during highway construction unearthed 11 specimens of fragmentary and unassociated avian fossils, which were identified by Rasmussen as including a small loon, a small gull-like species and five specimens of a gannet-like seabird, probably Morus loxostylus, a common species in the Miocene. All of these forms were already known from a site in Chesapeake Bay, Maryland. The finds suggests that the Delaware site was the near-shore area of a large bay at the time of deposition.[35]

Rasmussen was also involved in a review of fossil birds from Miocene and Pliocene deposits in North Carolina. Finds included an early Miocene loon Colymboides minutus, various ducks, a crested tern closely resembling the modern royal tern Sterna maxima, and a member of the crow genus, one of the few fossil passerine birds from that period. The review found that fossil birds from this period generally closely resemble a modern species or genus, and those that do not can usually be placed in a modern family with a fair degree of confidence.[36]

Birds of South Asia

In 1992, Rasmussen took the position of assistant to S. Dillon Ripley, the former secretary of the Smithsonian, who was planning to produce a definitive guide to the birds of South Asia. When he became ill shortly after beginning the project, Rasmussen took over the project, and with artist John C. Anderton, produced Birds of South Asia: The Ripley Guide, a two-volume bird guide for the Indian subcontinent which was the first field guide for the area to include sonograms. Volume 1 contains the field guide with over 3400 illustrations in 180 plates, and more than 1450 colour maps. Volume 2 (Attributes and Status) gives specimen measurements, data about identification, status, distribution and habits. Vocalizations are described from recordings, and there are over 1000 sonagrams.[37]

1508 species that have occurred in India, Bangladesh, Pakistan, Nepal, Bhutan, Maldives, the Chagos archipelago and Afghanistan are covered, including 85 hypothetical and 67 'possible' species, which are given only short accounts. Notable aspects of Birds of South Asia are its distribution evidence-base – the book's authors based their distributional information almost completely on museum specimens – and its taxonomic approach, involving a large number of species-level splits. Its geographical range was also greater than that of older works, notably in the inclusion of Afghanistan.[37]

Many allopatric forms previously regarded as conspecific are treated by Rasmussen and Anderton as full species. Most of these had previously been proposed elsewhere, but the book introduced a number of innovations of its own. Experts on Asian birds, Nigel Collar and John Pilgrim, in 2008 analysed Rasmussen and Anderton's proposed changes, indicating which had previously been proposed by other authors, and which were novel, and required further justification.[38]

Although reviews in the birding and ornithological press have often been favourable,[39][40] there have been criticisms. Peter Kennerley, author and Asian bird expert,[41] considered that some of the illustrations are small and garish or technically inaccurate. He also believes that the over-reliance on sometimes very old museum specimens and dismissal of the wealth of observational data filed by amateur travelling birders is a mistake, and states that many of the taxonomic decisions appear to be random choices, unsupported by published research.[42]

Apart from the Meinertzhagen fraud, which is discussed in the next section, and the death of S. Dillon Ripley, other problems in the production of Birds of South Asia included the loss of the main map database during a trip to Burma, and poorly prepared specimen skins. There were also difficulties reconciling sources, delays in producing illustrations and maps, and in obtaining reliable data for "difficult" areas like Assam, Arunachal Pradesh, Bangladesh and Afghanistan. The Andaman and Nicobar Islands also presented serious challenges with regard to the status and taxonomy of their avifaunas.[43]

Rasmussen considered in a 2005 paper whether the revised taxonomy of the book, with its many species splits, had significant conservation implications, but felt that the effect on species richness in South Asia was limited, and would have only a moderate conservation impact, increasing the number of potentially threatened species in the region from 6% of the total avifauna to about 7%.[44]

Meinertzhagen fraud

Hill blue-flycatcher (formerly large blue flycatcher), a species with false Meinertzhagen records[43]

Rasmussen revealed the true extent of the major fraud perpetrated by the eminent British soldier, ornithologist and expert on bird lice, Colonel Richard Meinertzhagen. Meinertzhagen, who died in 1967, was the author of numerous taxonomic and other works on birds, and possessed a vast collection of bird and bird lice specimens; he was considered to be one of Britain's greatest ornithologists. However, British ornithologist Alan Knox had analysed Meinertzhagen's bird collection at the Walter Rothschild Zoological Museum in Tring, UK in the early 1990s, and uncovered significant fraud involving theft of specimens from museums and falsification of the accompanying documentation.[45]

When researching for Birds of South Asia, Rasmussen examined tens of thousands of bird specimens, since the late S. Dillon Ripley had strongly favoured the use of museum specimens to determine which birds to include. With Robert Prys-Jones of the Natural History Museum, she showed that the decades-old Meinertzhagen fraud was far more extensive than first thought.[46] Many of the 20,000 bird specimens in his collection had been relabelled with regard to where they were collected, and sometimes also remounted. The false documentation delayed the rediscovery of the forest owlet, since previous searches had relied on Meinertzagen's faked records. Rasmussen's successful expedition ignored these and looked in the areas identified by the remaining genuine specimens.[47]

Meinertzhargen had been banned from the Natural History Museum's Bird Room for 18 months for unauthorised removal of specimens, and suspicions that he was stealing specimens and library material were documented by staff for over 30 years, twice reaching the verge of prosecution.[47]

Falsified records identified by Rasmussen and Prys-Jones included high-altitude occurrences of coral-billed scimitar-babbler Pomatorhinus ferruginosus, out-of-range Kashmir flycatcher Ficedula subrubra and Himalayan winter records of ferruginous flycatcher Muscicapa ferruginea and large blue flycatcher Cyornis magnirostris (now hill blue-flycatcher C. banyumas).[43][48] However, some records such as those for Afghan snowfinch Montifringilla theresae, a species Meinertzhagen described, appear to be genuine.[43]


  1. ^ a b Seabrook, John (29 May 2006). "Ruffled Feathers" (PDF). New Yorker. Archived from the original (PDF) on 13 June 2006. Retrieved 8 May 2008.
  2. ^ Rasmussen, Pamela C (1990). Geographic variation and evolutionary history of blue-eyed shags of South America (Phalacrocoracidae: Phalacrocorax [Notocarbo]) (Thesis). University of Kansas: Systematics and Ecology.
  3. ^ Johnston, Richard F. (1995). "Ornithology at the University of Kansas". In Davis, Jr., W. E.; Jackson, J. A. (eds.). Contributions to the history of North American ornithology. II. Cambridge, Massachusetts, US: Nuttall Ornithological Club. p. 104. ISBN 978-1-877973-40-6.
  4. ^ a b "Pamela Rasmussen". Curator Profiles. Michigan State University. Archived from the original on June 9, 2010. Retrieved 6 February 2008.
  5. ^ "Michael D. Gottfried". Curator Profiles. Michigan State University. Archived from the original on June 9, 2010. Retrieved 18 June 2008.
  6. ^ Rasmussen, P. C. (1986). "Reevaluation of cheek patterns of juvenal-plurnaged blue-eyed and king shags". Condor. 88 (3): 393–95. doi:10.2307/1368895. JSTOR 1368895.
  7. ^ Rasmussen, P. C. (1988). "Variation in the juvenal plumage of the red-legged shag (Phalacrocorax gaimardi) and notes on behavior of juveniles" (PDF). Wilson Bulletin. 100 (4): 535–44. JSTOR 4162642.
  8. ^ Rasmussen, P. C.; Hurnphrey, P. S. (1988). "Wing-spreading in Chilean blue-eyed shags (Phalacrocorax atriceps)". Wilson Bulletin. 100 (1): 140–44. JSTOR 4162533.
  9. ^ Rasmussen, P. C. (1989). "Post-landing displays of Chilean blue-eyed shags at a cliff-nesting colony". Bird Behaviour. 8: 51–54. doi:10.3727/015613888791871322.
  10. ^ Rasmussen, P. C. (1991). "Relationships between South American king and blue-eyed shags" (PDF). The Condor. 93 (4): 825–39. doi:10.2307/3247717. JSTOR 3247717.
  11. ^ Humphrey, P. S.; Rasmussen, P. C.; Lopez, N. (1988). "Fish surface activity and pursuit-plunging by olivaceous cormorants" (PDF). Wilson Bulletin. 100 (2): 327–28.
  12. ^ Rasmussen, Pamela (1998). "A new species of Scops-owl from Great Nicobar Island". Bulletin of the British Ornithologists' Club. 118: 141–151, pl. 3.
  13. ^ Lambert, Frank R; Rasmussen, Pamela (1998). "Sangihe Scops Owl Otus collari, sp. nov" (PDF). Bulletin of the British Ornithologists' Club. 118: 207–217. Archived from the original (PDF) on 2013-07-20. Retrieved 2013-11-24.
  14. ^ Rasmussen, P.C. (1999). "A New Species of Hawk-owl Ninox from North Sulawesi, Indonesia". Wilson Bulletin. 111 (4): 457–464.
  15. ^ Rasmussen, Pamela C.; Round, Philip D.; Dickinson, Edward C.; Rozendaal, F.G. (Apr 2000). "A new bush-warbler (Sylviidae, Bradypterus) from Taiwan". The Auk. 117 (2): 279. doi:10.1642/0004-8038(2000)117[0279:ANBWSB]2.0.CO;2.
  16. ^ Rasmussen, P. C.; Ishtiaq, F. (1999). "Vocalizations and Behaviour of Forest Spotted Owlet Athene blewitti" (PDF). Forktail. 15: 61–66. Archived from the original (PDF) on 2011-11-19.
  17. ^ Rasmussen, P. C.; King, B. F. (1998). "The rediscovery of the Forest Owlet Athene (Heteroglaux) blewitti" (PDF). Forktail. 14: 53–55. Archived from the original (PDF) on 2012-02-09.
  18. ^ Ripley, S. D. (1976). "Reconsideration of Athene blewitti (Hume)". Journal of the Bombay Natural History Society. 73: 1–4.
  19. ^ a b Rasmussen, Pamela C. (1998). "Rediscovery of an Indian enigma: the Forest Owlet". Bulletin of the Oriental Bird Club. 27.
  20. ^ Rasmussen, P. C.; Wardill, J. C; Lambert, F. R.; Riley, J. (2000). "On the specific status of the Sangihe White-eye Zosterops nehrkorni, and the taxonomy of the Black-crowned White-eye Z. atrifrons complex" (PDF). Forktail. 16: 69–80. Archived from the original (PDF) on 2011-06-10.
  21. ^ Warakagoda, Deepal H; Rasmussen, Pamela C. (2004). "A new species of scops-owl from Sri Lanka" (PDF). Bulletin of the British Ornithologists' Club. 124 (2): 85–105. Archived from the original (PDF) on 2014-02-07. Retrieved 2013-11-24.
  22. ^ Hennache, A.; Rasmussen, P.; Lucchini, V.; Rimoldi, S.; Randi, E. (2003). "Hybrid origin of the Imperial Pheasant Lophura imperialis (Delacour and Jabouille, 1924) demonstrated by morphology, hybrid experiments, and DNA analyses". Biological Journal of the Linnean Society. 80 (4): 573–600. doi:10.1111/j.1095-8312.2003.00251.x.
  23. ^ a b Indrawan, Mochamad; Rasmussen, Pamela C.; Sunarto (March 2008). "A New White-Eye (Zosterops) from the Togian Islands, Sulawesi, Indonesia". The Wilson Journal of Ornithology. 120 (1): 1–9. doi:10.1676/06-051.1.
  24. ^ Oaks, J. L; Gilbert, M; Virani, M.Z; Watson, R. T; Meteyer, C. U; Rideout, B. A; Shivaprasad, H. L; Ahmed, S; Chaudhry, M, J; Arshad, M; Mahmood, S; Ali, A; Khan, A. Al (2004). "Diclofenac residues as the cause of vulture population decline in Pakistan" (PDF). Nature. 427 (6975): 630–3. doi:10.1038/nature02317. PMID 14745453. Archived from the original (PDF) on 2006-09-08.CS1 maint: Multiple names: authors list (link)
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  26. ^ Johnson, Jeff A.; Lerner, Heather R. L.; Rasmussen, Pamela C.; Mindell, David P. (2006). "Systematics within Gyps vultures: a clade at risk". BMC Evolutionary Biology. 6: 65. doi:10.1186/1471-2148-6-65. PMC 1569873.
  27. ^ Orme, C. David L.; Davies, Richard G; Burgess, Malcolm; Eigenbrod, Felix;; Pickup, Nicola; Olson, Valerie A; Webster, Andrea J.; Ding, Tzung-Su; Rasmussen, Pamela C; Ridgely, Robert S; Stattersfield, Ali J.; Bennett, Peter M; Blackburn, Tim M; Gaston, Kevin J; Owens, Ian P. F. (August 2005). "Global hotspots of species richness are not congruent with endemism or threat" (PDF). Nature. 436 (7053): 1016–1019. doi:10.1038/nature03850. PMID 16107848.
  28. ^ Stevens, G. C. (1989). "The latitudinal gradient in geographical range: How so many species co-exist in the tropics" (PDF). American Naturalist. 133 (2): 240–256. doi:10.1086/284913. Archived from the original (PDF) on 2014-08-09.
  29. ^ Orme, C David L; Davies, Richard G; Olson, Valerie A; Thomas, Gavin H; Ding, Tzung-Su; Rasmussen, Pamela C; Ridgely, Robert S; Stattersfield, Ali J; Bennett, Peter M; Blackburn, Tim M; Owens, Ian P. F; Gaston, Kevin J (2006). "Global Patterns of Geographic Range Size in Birds". PLoS Biol. 4 (7): e208. doi:10.1371/journal.pbio.0040208. PMC 1479698. PMID 16774453.
  30. ^ Davies, Richard G; Orme, C David L; Olson, Valerie A; Thomas, Gavin H; Ross, Simon G; Ding, Tzung-Su; Rasmussen, Pamela C; Stattersfield, Ali J; Bennett, Peter M; Blackburn, Tim M; Owens, Ian P. F; Gaston, Kevin J (September 2006). "Human impacts and the global distribution of extinction risk" (PDF). Proceedings of the Royal Society B. 273 (1598): 2127–2133. doi:10.1098/rspb.2006.3551. PMC 1635517. PMID 16901831. Archived from the original (PDF) on 2008-09-20.
  31. ^ Grenyer, Richard; Orme, C David L; Jackson, Sarah F; Thomas, Gavin H; Davies, Richard G; Davies, T Jonathan; Jones, Kate E; Olson, Valerie A; Ridgely, Robert S; Ding, Tzung-Su; Bennett, Peter M; Blackburn, Tim M; Owens, Ian P. F; Gaston, Kevin J; Gittleman, John L; Owens, Ian. P. F. (November 2006). "Global distribution and conservation of rare and threatened vertebrates". Nature. 444 (7115): 93–6. doi:10.1038/nature05237. PMID 17080090.
  32. ^ Storch, David; Davies, Richard G; Zajícek, Samuel; Orme, C David L; Olson, Valerie A; Thomas, Gavin H; Ding, Tzung-Su; Rasmussen, Pamela C; Ridgely, Robert S; Bennett, Peter M; Blackburn, Tim M; Owens, Ian P. F; Gaston, Kevin J (December 2006). "Energy, range dynamics and global species richness patterns: reconciling mid-domain effects and environmental determinants of avian diversity". Ecology Letters. 9 (12): 1308–20. doi:10.1111/j.1461-0248.2006.00984.x. PMID 17118005.
  33. ^ Davies, Richard G; Orme, C David L; Storch, David; Olson, Valerie A; Thomas, Gavin H; Ross, Simon G; Ding, Tzung-Su; Rasmussen, Pamela C; Lennon, Jack J; Bennett, Peter M; Owens, Ian P. F; Blackburn, Tim M; Gaston, Kevin J (2007). "Increased genetic diversity as a defence against parasites is undermined by social parasites: Microdon mutabilis hoverflies infesting Formica lemani ant colonies" (PDF). Proceedings of the Royal Society B. 274 (1606): 1189–1197. doi:10.1098/rspb.2006.0061. PMC 1679886. PMID 17035169.
  34. ^ Gaston, Kevin J; Davies, Richard G; Orme, C David L; Olson, Valerie A; Thomas, Gavin H; Ding, Tzung-Su; Rasmussen, Pamela C; Lennon, Jack J; Bennett, Peter M; Owens, Ian P. F; Blackburn, Tim M (July 2007). "Spatial turnover in the global avifauna". Proceedings of the Royal Society B. 274 (1618): 1567–74. doi:10.1098/rspb.2007.0236. PMC 2169276. PMID 17472910.
  35. ^ Rasmussen, Pamela C. (1998) "Early Miocene avifauna from the Pollack Farm site, Delaware Archived 2009-02-16 at the Wayback Machine" (PDF) in Benson, Richard N. (editor) (1998) Geology and paleontology of the lower Miocene Pollack Farm fossil site, Delaware Archived 2009-02-16 at the Wayback Machine Delaware Geological Survey Special Publication no. 21, State of Delaware & University of Delaware, 144–146
  36. ^ "Miocene and Pliocene birds from the Lee Creek Mine, North Carolina" in Ray, C. E. & Bohaska, D. J. (2001). "Geology and Paleontology of the Lee Creek Mine, North Carolina, III." Smithsonian Contributions to Paleobiology, 90. Smithsonian Institution Press, Washington D.C. 233–365.
  37. ^ a b Rasmussen, Pamela C; Anderton, John C. (2005): Birds of South Asia: The Ripley Guide. Lynx Edicions, Barcelona. ISBN 84-87334-67-9
  38. ^ Collar, Nigel J; Pilgrim John, D. (2008). "Taxonomic Update: Species-level changes proposed for Asian birds, 2005–2006". BirdingASIA. 8: 14–30.
  39. ^ Bishop, K. David; Myers, Susan D. (2006). "Book review: Birds of South Asia: The Ripley Guide Volume i: field guide, Volume ii: attributes and status" (PDF). Emu. 106: 87–91. doi:10.1071/MUv106n1_BR.
  40. ^ Dickinson, Edward C. (2006). "Birds of South Asia: The Ripley Guide". The Auk. 123 (3): 916–918. doi:10.1642/0004-8038(2006)123[916:BOSATR]2.0.CO;2.
  41. ^ Kennerley, Peter; Pearson, David (2008). Reed and Bush Warblers. Christopher Helm. ISBN 978-0-7136-6022-7.
  42. ^ "The Ripley Guide, Volumes 1 and 2". Reviews. Retrieved 2008-04-18.
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  44. ^ Rasmussen, P.C. (2005). "Biogeographic and conservation implications of revised species limits and distributions of South Asian birds" (PDF). Zoologische Mededelingen. 79 (3): 137–146.
  45. ^ Knox, Alan G. (1993). "Richard Meinertzhagen—a case of fraud examined". Ibis. 135 (3): 320–325. doi:10.1111/j.1474-919X.1993.tb02851.x.
  46. ^ Rasmussen, P. C.; Prŷs-Jones, R. P. (2003). Collar, N. J.; Fisher, C. T.; Feare, C. J. (eds.). "History vs mystery: the reliability of museum specimen data". Bulletin of the British Ornithologists' Club. 123A (Why Museums Matter: Avian Archives in an Age of Extinction): 66–94.
  47. ^ a b "Bird collection fraud". News 17 November 2005. Natural History Museum. Archived from the original on 10 April 2008. Retrieved 23 April 2008.
  48. ^ BirdLife International 2004. Cyornis banyumas. 2006 IUCN Red List of Threatened Species. Downloaded on 24 April 2008
1990 in paleontology

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 1990.

2001 in paleontology

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2001.

2005 in science

The year 2005 in science and technology involved some significant events.

Adelaide's warbler

Adelaide's warbler (Setophaga adelaidae) is a bird endemic to the archipelago of Puerto Rico belonging to the genus Setophaga of the family Parulidae. The species is named after Adelaide Swift, daughter of Robert Swift, the person who captured the first specimen.

Bahama warbler

The Bahama warbler (Setophaga flavescens) is a species of bird in the Parulidae family. It is endemic to The Bahamas.

The taxon was formerly lumped with the yellow-throated warbler (Setophaga dominica), until the Bahama warbler was elevated to full species in 2011.Its natural habitat is pine forest on Grand Bahama, Little Abaco and Great Abaco islands.

Banggai crow

The Banggai crow (Corvus unicolor) is a member of the crow family from Banggai regency in the province of Central Sulawesi in Indonesia. It is listed as critically endangered by IUCN. It was feared extinct, but was finally rediscovered during surveys on Peleng Island off the southeast coast of Sulawesi by Indonesian ornithologist Mochamad Indrawan in 2007 and 2008.

It was sometimes considered a subspecies of the slender-billed crow, but it is actually rather distinct from this bird, resembling an entirely black piping crow overall. The Banggai crow is a small crow, some 39 cm long and completely black with a pale iris and a short tail.For more than a century, it was known from only two specimens taken from an unknown island in the Banggai Archipelago - probably in 1884/1885. Visits to the archipelago in 1991 and 1996 yielded no unequivocal records of the species, leading some to believe it was extinct. During a survey conducted between 2007 and 2008 and partially financed by the Zoological Society for the Conservation of Species and Populations (Germany), it was repeatedly seen on Peleng Island and Indonesian ornithologist Mochamad Indrawan caught and photographed two individuals. The validity of the crows on Peleng was not recognized by BirdLife International in its 2009 Red List. Confirmation of the identity based on two specimens from Peleng was made by Pamela C. Rasmussen of the American Museum of Natural History in October 2009.

The total population is estimated at approximately 500 mature individuals, living in mountain forest at altitudes above 500 m. The decline of the Banggai crow is thought to be primarily due to habitat loss and degradation through agriculture and extraction.

This bird remained a complete enigma for a long time. Listed as Vulnerable in the 1994 IUCN Red List, it was changed to Endangered in 2000. In 2006, the status was considered as Possibly Extinct. This proved to be incorrect and the status was corrected to Critically Endangered in the 2007 Red List.

Birds of South Asia. The Ripley Guide

Birds of South Asia. The Ripley Guide by Pamela C. Rasmussen and John C. Anderton is a two-volume ornithological handbook, covering the birds of South Asia, published in 2005 (second edition in 2012) by the Smithsonian Institution and Lynx Edicions. The geographical scope of the book covers India, Bangladesh, Pakistan, Sri Lanka, Nepal, Bhutan, Maldives, the Chagos archipelago and Afghanistan (the latter country had been excluded from previous works covering this region). In total, 1508 species are covered (this figure includes 85 hypothetical and 67 'possible' species, which are given only shorter accounts). Two notable aspects of Birds of South Asia are its distribution evidence-base — the book's authors based their distributional information almost completely on museum specimens — and its taxonomic approach, involving a large number of species-level splits.

Cinnabar boobook

The cinnabar boobook (Ninox ios) also known as the cinnabar hawk owl, is a hawk owl endemic to the island of Sulawesi, Indonesia. It was described as a new species to science by American ornithologist Pamela C. Rasmussen in 1999 based on a single specimen collected by Frank Rozendaal from Bogani Nani Wartabone National Park on Minahassa Peninsula, northern Sulawesi, in 1985. Subsequently, it has also been observed in Lore Lindu National Park in central Sulawesi, greatly expanding the known habitat range.

The cinnabar boobook is small (total length 22 cm or 8.5 in) and has a relatively long tail and narrow pointed wings. The four known records of the species indicate it is a nocturnal forest dwelling species living at mid-altitudes (1,100 to 1,700 m (3,600 to 5,600 ft)). Otherwise very little is known of its habits. Based on morphological similarities with owlet-nightjars, Rasmussen suggests the cinnabar boobook may be an insectivore and prey on invertebrates in flight.

Common gallinule

The common gallinule (Gallinula galeata) is a bird in the family Rallidae. It was split from the common moorhen by the American Ornithologists' Union in July 2011. It lives around well-vegetated marshes, ponds, canals, and other wetlands in the Americas. The species is not found in the polar regions or many tropical rainforests. Elsewhere, the common gallinule is likely the most commonly seen rail species in much of North America, excepting the American coot in some regions.

Forest owlet

The forest owlet (Athene blewitti) is an endangered owl that is endemic to the forests of central India. The species belongs to the typical owls family, Strigidae. First described in 1873, it was not seen after 1884 and considered extinct until it was rediscovered 113 years later in 1997 by Pamela Rasmussen. Searches for the species in the locality given on the label of the last collected specimen failed and it was discovered that the specimen had been stolen from the British Museum by Richard Meinertzhagen and resubmitted with a label bearing false locality information. It is known from a small number of localities and the populations are very low within the fragmented and shrinking forests of central India.

Since its rediscovery in 1997, this species has been found at a number of new locations, several of which appear to hold significant populations. The population estimate has therefore been revised upwards, leading to its reclassification from Critically Endangered to Endangered. Nevertheless, the total known population remains very small and fragmented and is inferred to be declining as a result of the loss and

degradation of its deciduous forest habitat.

Lesser yellownape

The lesser yellownape (Picus chlorolophus) is a type of woodpecker which is a widespread and often common breeder in tropical and sub-tropical Asia, primarily the Indian subcontinent and Southeast Asia. It ranges from India, Bhutan, Nepal, Bangladesh and Sri Lanka eastwards to Thailand, Burma, Cambodia, Laos, Indonesia, Malaysia and Vietnam. Much of the scientific knowledge gathered about this species is sourced from formal studies in various parts of India.

Nicobar scops owl

The Nicobar scops owl (Otus alius) is a species of owl in the family Strigidae.

It is endemic to the Nicobar Islands, India, in particular Great Nicobar Island, but it may also occur on Little Nicobar Island.Its natural habitat is tropical moist lowland forests. It has an uncertain status but is thought to be rare or endangered. It was originally discovered by Pamela C. Rasmussen in 1998. Very little is known about this species, but individuals have been reported to consume spiders, beetles, and geckos.

Nilgiri blue robin

The Nilgiri blue robin (Sholicola major), also known as Nilgiri shortwing, white-bellied shortwing, Nilgiri sholakili or rufous-bellied shortwing is a species of passerine bird in the family Muscicapidae endemic to the Shola forests of the higher hills of southern India, mainly north of the Palghat Gap. This small bird is found on the forest floor and undergrowth of dense forest patches sheltered in the valleys of montane grassland, a restricted and threatened habitat.

The white-bellied blue robin was formerly considered conspecific with this species but in 2005 the two taxons were split by Pamela C. Rasmussen, a treatment that is followed by some authorities. Their genus remained uncertain until a 2017 molecular phylogenetic study found that these two south Indian species formed a sister group to a clade containing the genera Eumyias, Niltava and Cyornis. A new genus Sholicola was therefore erected for these two species.

Pamela (name)

Pamela is a feminine given name. Pamela is often abbreviated to Pam. Pamela is infrequently used as a surname.

Sangihe white-eye

The Sangihe white-eye (Zosterops nehrkorni) is a species of bird in the white-eye family. It is endemic to Sangihe, Indonesia.

Its natural habitat is subtropical or tropical moist montane forests. It is threatened by habitat loss.

It was formerly considered conspecific with black-crowned white-eye (Zosterops atrifrons), but work by Pamela C. Rasmussen and her colleagues showed that it is a separate species. The same research also confirmed the specific status of the Seram white-eye, Zosterops stalkeri.

Seram white-eye

The Seram white-eye (Zosterops stalkeri) is a small passerine bird in the white-eye family. It is an endemic resident breeder in open woodland in Seram, Indonesia.

It was formerly considered conspecific with black-fronted white-eye, Zosterops minor, but work by Pamela C. Rasmussen and her colleagues showed that it is a separate species. The same research also confirmed the specific status of the Sangihe white-eye, Zosterops nehrkorni.

Compared to related taxa, the bill of Seram white-eye is paler, deeper, and broader at the base. Its eye-ring is narrow and broken at the front. The crown and sides of the head are black and the upperparts are dark bronze. The rump is a distinctive yellow-bronze. The sides of the breast and flanks are greyish-white, the undertail-coverts are orange-yellow, the thighs are whitish, and the uppertail is brownish-black.

The sexes are similar, but immatures have the throat greener and more diffuse, with more black mixed into the chin feathers.

Its song also differs from that of related species.

Though mainly insectivorous, the Seram white-eye will also eat nectar and fruits of various kinds.


The order Suliformes (dubbed "Phalacrocoraciformes" by Christidis & Boles 2008) is an order recognised by the International Ornithologist's Union. In regard to the recent evidence that the traditional Pelecaniformes is polyphyletic, it has been suggested that the group be split up to reflect the true evolutionary relationships.

Transvolcanic jay

The Transvolcanic jay (Aphelocoma ultramarina) is a medium-large (~120 g) passerine bird similar in size to most other jays, with a blue head, blue-gray mantle, blue wings and tail, gray breast and underparts. The sexes are morphologically similar, and juveniles differ only in having less blue coloration. The iris is brown and legs are black. It is most readily distinguished by the plain (unstreaked) throat and breast, and the mantle contrasting less with the head and wings.

White-bellied blue robin

The white-bellied blue robin or white-bellied sholakili (Sholicola albiventris) refers to a kind of bird in the family Muscicapidae endemic to the Shola forests of the higher hills of southern India. The Nilgiri blue robin and this species were once considered separate species, later lumped as sub-species of a single species (major) and elevated again to full species in 2005 by Pamela C. Rasmussen. The species was earlier thought to be related to the shortwings and placed in the genus Brachypteryx and later moved to Myiomela since species in the genus Brachypteryx shows marked sexual dimorphism. In 2017, a study found that this is a sister group of the flycatchers in the genera Niltava, Cyornis and Eumyias among others. It was then placed in newly erected genus Sholicola. This small bird is found on the forest floor and undergrowth of dense forest patches sheltered in the valleys of montane grassland, a restricted and threatened habitat.


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