Ornithomimus

Ornithomimus (/ˌɔːrnɪθəˈmaɪməs, -θoʊ-/;[2] "bird mimic") is a genus of ornithomimid dinosaurs from the Late Cretaceous Period of what is now North America. Ornithomimus was a swift bipedal theropod which fossil evidence indicates was covered in feathers, equipped with a small toothless beak that may indicate an omnivorous diet. It is usually classified into two species: the type species, Ornithomimus velox, and a referred species, Ornithomimus edmontonicus. O. velox was named in 1890 by Othniel Charles Marsh on the basis of a foot and partial hand from the late Maastrichtian-age Denver Formation of Colorado, United States. Another seventeen species have been named since, though most of them have subsequently been assigned to new genera or shown to be not directly related to Ornithomimus velox. The best material of species still considered part of the genus has been found in Alberta, Canada, representing the species O. edmontonicus, known from several skeletons from the early Maastrichtian Horseshoe Canyon Formation. Additional species and specimens from other formations are sometimes classified as Ornithomimus, such as Ornithomimus samueli (alternately classified in the genera Dromiceiomimus or Struthiomimus) from the earlier, Campanian-age Dinosaur Park Formation of Alberta.

Ornithomimus
Temporal range: Late Cretaceous, 76.5–66.5 Ma[1]
OrnithomimusROM
Mounted O. edmontonicus skeleton, Royal Ontario Museum
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Order: Saurischia
Suborder: Theropoda
Clade: Ornithomimosauria
Family: Ornithomimidae
Genus: Ornithomimus
Marsh, 1890
Type species
Ornithomimus velox
Marsh, 1890
Other species

O. edmontonicus
Sternberg, 1933
O. samueli?
(Parks, 1928)

Description

Ornithomimus edmontonicus
Specimen of Ornithomimus edmontonicus found in 1995 with quill knobs, Royal Tyrrell Museum

Like other ornithomimids, species of Ornithomimus are characterized by feet with three weight-bearing toes, long slender arms, and long necks with birdlike, elongated, toothless, beaked skulls. They were bipedal and superficially resembled ostriches. They would have been swift runners. They had very long limbs, hollow bones, and large brains and eyes. The brains of ornithomimids in general were large for non-avialan dinosaurs, but this may not necessarily be a sign of greater intelligence; some paleontologists think that the enlarged portions of the brain were dedicated to kinesthetic coordination.[3] The bones of the hands are remarkably sloth-like in appearance, which led Henry Fairfield Osborn to suggest that they were used to hook branches during feeding.

Ornithomimus differ from other ornithomimids, such as Struthiomimus, in having shorter torsos, long slender forearms, very slender, straight hand and foot claws and in having hand bones (metacarpals) and fingers of similar lengths.[4]

OrnithomimusSizeComparison
Size of the two valid species.

The two Ornithomimus species today seen as possibly valid differ in size. In 2010 Gregory S. Paul estimated the length of O. edmontonicus at 3.8 m (12 ft), its weight at 170 kilograms (370 lb). One of its specimens, CMN 12228, preserves a femur (thigh bone) 46.8 centimetres (18.4 in) long. O. velox, the type species of Ornithomimus, is based on material of a smaller animal. Whereas the holotype of O. edmontonicus, CMN 8632, preserves a second metacarpal eighty-four millimetres long, the same element with O. velox measures only fifty-three millimetres.

Feathers and skin

Ornithomimus, like many dinosaurs, was long thought to have been scaly. However, beginning in 1995, several specimens of Ornithomimus have been found preserving evidence of feathers.

In 1995, 2008 and 2009, three Ornithomimus edmontonicus specimens with evidence of feathers were found; two adults with carbonized traces on the lower arm, indicating the former presence of pennaceous feather shafts, and a juvenile with impressions of feathers, of which were up to five centimetres in length, in the form of hair-like filaments covering the rump, legs and neck was also discovered. The fact that the feather imprints were found in sandstone, previously thought to not be able to support such impressions, raised the possibility of finding similar structures with more careful preparation of future specimens. A study describing the fossils in 2012 concluded that O. edmontonicus was covered in plumaceous feathers at all growth stages, and that only adults had pennaceous wing-like structures, suggesting that wings may have evolved for mating displays.[5] In 2014, Christian Foth and others argued that the evidence was insufficient to conclude that the forelimb feathers of Ornithomimus were necessarily pennaceous, citing the fact that the monofilamentous wing feathers in cassowaries would likely leave similar traces.[6]

"Ornithomimus" sp. by Tom Parker
Life restoration of the plumage pattern suggested by specimens preserving feathers and skin

A fourth feathered specimen of Ornithomimus, this time from the lower portion of the Dinosaur Park Formation, was described in October, 2015 by Aaron van der Reest, Alex Wolfe, and Phil Currie. It was the first Ornithomimus specimen to preserve the feathers along the tail. The feathers, though crushed and distorted, bore numerous similarities with those of the modern ostrich, both in their structure and distribution on the body. Skin impressions were also preserved in the 2015 specimen, which indicated that from mid-thigh to the feet, there was bare skin devoid of scales, and that a flap of skin connect the upper thigh to the torso. This latter structure is similar to that found in modern birds, including ostriches, but was positioned higher above the knee in Ornithomimus than in birds.[7]

History of discovery

First species named

Ornithomimus velox
Holotype material of O. velox

The history of Ornithomimus classification, and the classification of ornithomimids in general, has been complicated. The type species, Ornithomimus velox, was first named by O.C. Marsh in 1890, based on syntypes YPM 542 and YPM 548, a partial hindlimb and forelimb found on 30 June 1889 by George Lyman Cannon in the Denver Formation of Colorado. The generic name means "bird mimic", derived from Greek ὄρνις, ornis, "bird", and μῖμος, mimos, "mimic", in reference to the bird-like foot. The specific name means "swift" in Latin.[8] Simultaneously, Marsh named two other species: Ornithomimus tenuis, based on specimen USNM 5814, and Ornithomimus grandis. Both consist of fragmentary fossils found by John Bell Hatcher in Montana of which it is today understood they represent tyrannosauroid material. At first Marsh assumed Ornithomimus was an ornithopod but this changed when Hatcher found specimen USNM 4736, a partial ornithomimid skeleton, in Wyoming, which Marsh named Ornithomimus sedens in 1892. On that occasion also Ornithomimus minutus was created based on specimen YPM 1049, a metatarsus,[9] since recognized as belonging to the Alvarezsauridae.[10]

A sixth species, Ornithomimus altus, was named in 1902 by Lawrence Lambe, based on specimen CMN 930, hindlimbs found in 1901 in Alberta,[11] but this was in 1916 renamed to a separate genus, Struthiomimus, by Henry Fairfield Osborn.[12] In 1920 Charles Whitney Gilmore named Ornithomimus affinis for Dryosaurus grandis Lull 1911,[13] based on indeterminate material. In 1930 Loris Russell renamed Struthiomimus brevetertius Parks 1926 and Struthiomimus samueli Parks 1928 into Ornithomimus brevitertius and Ornithomimus samueli.[14] The same year Oliver Perry Hay renamed Aublysodon mirandus Leidy 1868 into Ornithomimus mirandus,[15] today seen as a nomen dubium. In 1933 William Arthur Parks created a Ornithomimus elegans,[16] today seen as either belonging to Chirostenotes or Elmisaurus. That same year, Gilmore named Ornithomimus asiaticus for material found in Inner Mongolia.[17]

Also in 1933, Charles Mortram Sternberg named the species Ornithomimus edmontonicus for a nearly complete skeleton from the Horseshoe Canyon Formation of Alberta, specimen CMN 8632.[18]

Reclassification by Dale Russell

Ornithomimid skull
Skull and neck of Ornithomimus sp. (RTMP 95.110.1)

At first it had been common to name each newly discovered ornithomimid as a species of Ornithomimus. In the sixties, this tendency was still strong as is shown by the fact that Oskar Kuhn renamed Megalosaurus lonzeensis Dollo 1903 from Belgium into Ornithomimus lonzeensis (today understood to be an abelisauroid claw),[19] and Dale Russell in 1967 renamed Struthiomimus currellii Parks 1933 and Struthiomimus ingens Parks 1933 into Ornithomimus currellii and Ornithomimus ingens.[20] At the same time it was usual that workers referred to the entire ornithomimid material as simply "Struthiomimus".[21] To solve this confusion by scientifically testing the separation between Ornithomimus and Struthiomimus, in 1972 Dale Russell published a morphometric study showing that statistical differences in some proportions could be used to distinguish the two. He concluded that Struthiomimus and Ornithomimus were valid genera. In the latter Russell recognised two species: the type species Ornithomimus velox and Ornithomimus edmontonicus even though he had trouble reliably distinguishing it from O. velox. Struthiomimus currellii he considered a younger synonym of Ornithomimus edmontonicus. However, Russell also interpreted the data as indicating that many specimens could not be referred to either Ornithomimus or Struthiomimus. Therefore, he created two new genera. The first was Archaeornithomimus to which Ornithomimus asiaticus and Ornithomimus affinis were assigned, becoming an Archaeornithomimus asiaticus and an Archaeornithomimus affinis. The second genus was Dromiceiomimus, meaning "Emu mimic" from the old generic name for the emu, Dromiceius. Russell assigned several former Ornithomimus species named during the 20th century, including O. brevitertius and O. ingens, to the new genus as Dromiceiomimus brevitertius. He renamed Ornithomimus samueli into a second Dromiceiomimus species: Dromiceiomimus samueli.[22]

Misassigned to Ornithomimus

Two tibiae from the Navesink Formation of New Jersey were named Coelosaurus antiquus ("antique hollow lizard") by Joseph Leidy in 1865. The tibiae were first attributed to Ornithomimus in 1979 by Donald Baird and John R. Horner as Ornithomimus antiquus.[23] Normally, this would have made Ornithomimus a junior synonym of Coelosaurus, but Baird and Horner discovered that the name "Coelosaurus" was preoccupied by a dubious taxon based on a single vertebra, named Coelosaurus by an anonymous author now known to be Richard Owen in 1854.[24] Baird referred several other specimens from New Jersey and Maryland to O. antiquus. Beginning in 1997, Robert M. Sullivan regarded O. velox and O. edmontonicus as junior synonyms of O. antiquus. Like Russell, he considered the former two species indistinguishable from each other, and noted that they both shared distinctive features with O. antiquus.[25] However, David Weishampel (2004) considered "C." antiquus to be indeterminate among ornithomimosaurs, and therefore a nomen dubium.[24] An SVP 2012 abstract agreed with Weishampel by noting that Coelosaurus differs from Gallimimus and Ornithomimus in the features of the tibiae.[26]

In 1988 Gregory S. Paul classified the species in genera Archaeornithomimus, Struthiomimus, Dromiceiomimus and Gallimimus in genus Ornithomimus.[27] This has found no acceptance among other workers and presently the name is not used by Paul himself.

Present interpretations

Ornithomimus skull
T scan of O. edmontonicus skull RTMP 1995.110.0001, with taphonomically deformed bones reconstructed on the right

Even after Russell's study, various researchers have found reasons to lump some or all of these species back into Ornithomimus in various combinations. In 2004, Peter Makovicky, Yoshitsugu Kobayashi and Phil Currie studied Russell's 1972 proportional statistics to re-analyze ornithomimid relationships in light of new specimens. They concluded that there was no justification to separate Dromiceiomimus from Ornithomimus, sinking Dromiceiomimus as a synonym of O. edmontonicus.[28] However, they did not include the type species of Ornithomimus, O. velox, in this analysis. The same team further supported the synonymy between Dromiceiomimus and O. edmontonicus in a 2006 lecture at the Society of Vertebrate Paleontology annual meeting,[29] and their opinion has been followed by most later authors.[30] Makovicky's team also considered Dromiceiomimus samueli to be a junior synonym of O. edmontonicus, though Longrich later suggested it may belong to a distinct, unnamed species from the Dinosaur Park Formation which have yet to be described.[30] Longrich called the species Ornithomimus samueli in a faunal list for the Dinosaur Park Formation.[31]

Apart from O. edmontonicus dating to the early Maastrichtian, two other species are presently considered to be possibly valid, both from the late Maastrichtian. O. sedens was named by Marsh in 1892 from partial remains found in the Lance Formation of Wyoming, only one year after the description of O. velox. Dale Russell, in his 1972 revision of ornithomimids, could not determine which genus it actually belonged to, though he speculated that it may be intermediate between Struthiomimus and Dromiceiomimus. In 1985 he considered it a species of Ornithomimus.[32] Although it has since been referred to mainly as Struthiomimus sedens, based on complete specimens from Montana (as well as some fragments from Alberta and Saskatchewan), these yet have to be described and compared to the O. sedens holotype.[30]

The other is the original type species: O. velox, at first known from very limited remains. Additional specimens referred to O. velox have been described from the Denver Formation and from the Ferris Formation of Wyoming.[33] One specimen attributed to O. velox (MNA P1 1762A) from the Kaiparowits Formation of Utah, was described in 1985.[32] Re-evaluation of this specimen by Lindsay Zanno and colleagues in 2010, however, cast doubt on its assignment to O. velox, and possibly even to Ornithomimus.[34] This conclusion was supported by a 2015 re-description of O. velox, which found that only the holotype specimen was confidently referable to that species. The authors of this study tentatively referred to the Kaiparowits specimen as Ornithomimus sp., along with all of the specimens from the Dinosaur Park Formation.[1]

Classification

Ornithomimus - claw
Claw bone

In 1890 Marsh assigned Ornithomimus to the Ornithomimosauria, a classification that is still common. Modern cladistic studies indicate a derived position in the ornithomimids; these however have only included O. edmontonicus in their analyses. The relationships between O. edmontonicus, O. velox and O. sedens have not been published.

The following cladogram is based on Xu et al., 2011:[35]

Ornithomimidae

Archaeornithomimus asiaticus

unnamed

Sinornithomimus dongi

unnamed

Anserimimus planinychus

Gallimimus bullatus

unnamed

Qiupalong henanensis

unnamed

Struthiomimus altus

Ornithomimus edmontonicus

Paleobiology

Dakotaraptor RPR
An Ornithomimus being restrained while preyed upon by Dakotaraptor

The diet of Ornithomimus is still debated. As theropods, ornithomimids might have been carnivorous but their body shape would also have been suited for a partly or largely herbivorous lifestyle. Suggested food includes insects, crustaceans, fruit, leaves, branches, eggs, and the meat of lizards and small mammals.[3]

Ornithomimus had legs that seem clearly suited for rapid locomotion, with the tibia about 20% longer than the femur. The large eye sockets suggest a keen visual sense, and also suggest the possibility that they were nocturnal.[36]

In a 2001 study conducted by Bruce Rothschild and other paleontologists, 178 foot bones referred to Ornithomimus were examined for signs of stress fracture, but none were found.[37]

See also

References

  1. ^ a b Claessens, L. & Mark A. Loewen, M.A. (2015). A redescription of Ornithomimus velour Marsh, 1890 (Dinosauria, Theropoda). Journal of Vertebrate Paleontology (advance online publication). doi:10.1080/02724634.2015.1034593
  2. ^ "Ornithomimus". Oxford Dictionaries. Oxford University Press. Retrieved 2016-01-21.
  3. ^ a b "Dromiceiomimus." In: Dodson, Peter & Britt, Brooks & Carpenter, Kenneth & Forster, Catherine A. & Gillette, David D. & Norell, Mark A. & Olshevsky, George & Parrish, J. Michael & Weishampel, David B. The Age of Dinosaurs. Publications International, LTD. p. 140. ISBN 0-7853-0443-6.
  4. ^ Makovicky, P.J., Kobayashi, Y., and Currie, P.J. (2004). "Ornithomimosauria." In Weishampel, D.B., Dodson, P., & Osmólska, H. (eds.), The Dinosauria (second edition). University of California Press, Berkeley: 137-150.
  5. ^ Zelenitsky, D. K.; Therrien, F.; Erickson, G. M.; Debuhr, C. L.; Kobayashi, Y.; Eberth, D. A.; Hadfield, F. (2012). "Feathered Non-Avian Dinosaurs from North America Provide Insight into Wing Origins". Science. 338 (6106): 510–514. Bibcode:2012Sci...338..510Z. doi:10.1126/science.1225376. PMID 23112330.
  6. ^ Christian Foth; Helmut Tischlinger; Oliver W. M. Rauhut (2014). "New specimen of Archaeopteryx provides insights into the evolution of pennaceous feathers". Nature. 511 (7507): 79–82. Bibcode:2014Natur.511...79F. doi:10.1038/nature13467. PMID 24990749.
  7. ^ Van Der Reest, Aaron J.; Wolfe, Alexander P.; Currie, Philip J. (2016). "A densely feathered ornithomimid (Dinosauria: Theropoda) from the Upper Cretaceous Dinosaur Park Formation, Alberta, Canada". Cretaceous Research. 58: 108–117. doi:10.1016/j.cretres.2015.10.004.
  8. ^ O.C. Marsh, 1890, "Description of new dinosaurian reptiles", The American Journal of Science, series 3 39: 81-86
  9. ^ O.C. Marsh, 1892, "Notice of new reptiles from the Laramie Formation", American Journal of Science 43: 449-453
  10. ^ Holtz, Thomas R. Jr. (2011) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Winter 2010 Appendix.
  11. ^ Lambe, L., 1902, "New genera and species from the Belly River Series (mid-Cretaceous)", Geological Survey of Canada Contributions to Canadian Palaeontology 3(2): 25-81
  12. ^ H.F. Osborn, 1916, "Skeletal adaptations of Ornitholestes, Struthiomimus, Tyrannosaurus", Bulletin of the American Museum of Natural History 35(43): 733-771
  13. ^ Gilmore, C.W., 1920, "Osteology of the carnivorous Dinosauria in the United States National Museum with special reference to the genera Antrodemus (Allosaurus) and Ceratosaurus, United States National Museum Bulletin 110: l-154
  14. ^ Russell, L.S., 1930, "Upper Cretaceous dinosaur faunas of North America", Proceedings of the American Philosophical Society, 69(4): 133-159
  15. ^ Hay, O.P., 1930, Second Bibliography and Catalogue of the Fossil Vertebrata of North America. Carnegie Institution of Washington. 390(II): 1-1074
  16. ^ Parks, W.A., 1933, "New species of dinosaurs and turtles from the Upper Cretaceous formations of Alberta", University of Toronto Studies, Geological Series, 34: 1-33
  17. ^ Gilmore, C.W., 1933, "On the dinosaurian fauna of the Iren Dabasu Formation", Bulletin of the American Museum of Natural History, 67: 23-78
  18. ^ Sternberg, C.M., 1933, "A new Ornithomimus with complete abdominal cuirass", The Canadian Field-Naturalist 47(5): 79-83
  19. ^ Kuhn, O., 1965, "Saurischia (Supplementum 1)". In: Fossilium Catalogus 1. Animalia. 109: 1-94
  20. ^ Russell, D.A. and Chamney, T.P., 1967, "Notes on the biostratigraphy of dinosaurian and microfossil faunas in the Edmonton Formation (Cretaceous), Alberta", National Museum of Canada Natural History Papers, 35: 1-22
  21. ^ Norman, D., 1985, The Illustrated Encyclopedia of Dinosaurs, Crescent Books, New York, p. 48
  22. ^ Russell, D. (1972). "Ostrich dinosaurs from the Late Cretaceous of western Canada." Canadian Journal of Earth Sciences, 9: 375-402.
  23. ^ Baird D., and Horner, J., 1979, "Cretaceous dinosaurs of North Carolina", Brimleyana 2: 1-28
  24. ^ a b Weishampel, D.B.(2004). "Another Look at the Dinosaurs of the East Coast of North America. En (Colectivo Arqueológico-Paleontológico Salense, Ed.). Archived 2012-09-04 at the Wayback Machine" Actas de las III Jornadas sobre Dinosaurios y su Entorno. 129-168. Salas de los Infantes, Burgos, España.
  25. ^ Sullivan, (1997). "A juvenile Ornithomimus antiquus (Dinosauria: Theropoda: Ornithomimosauria), from the Upper Cretaceous Kirtland Formation (De-na-zin Member), San Juan Basin, New Mexico." New Mexico Geological Society Guidebook, 48th Field Conference, Mesozoic Geology and Paleontology of the Four Corners Region. 249-254.
  26. ^ Brusatte, Choiniere, Benson, Carr and Norell, 2012. Theropod dinosaurs from the Late Cretaceous of Eastern North America: Anatomy, systematics, biogeography and new information from historic specimens. Journal of Vertebrate Paleontology. Program and Abstracts 2012, 70.
  27. ^ Paul, G.S., 1988, Predatory Dinosaurs of the World. Simon & Schuster: New York. 464 pp
  28. ^ Makovicky, Kobayashi and Currie (2004). "Ornithomimosauria." In Weishampel, Dodson and Osmolska (eds.), The Dinosauria Second Edition. University of California Press. 861 pp.
  29. ^ Kobayashi, Makovicky and Currie (2006). "Ornithomimids (Theropoda: Dinosauria) from the Late Cretaceous of Alberta, Canada." Journal of Vertebrate Paleontology, 26(3): 86A.
  30. ^ a b c Longrich, N. (2008). "A new, large ornithomimid from the Cretaceous Dinosaur Park Formation of Alberta, Canada: Implications for the study of dissociated dinosaur remains." Palaeontology, 51(4): 983-997.
  31. ^ Longrich, N. R. (2014). "The horned dinosaurs Pentaceratops and Kosmoceratops from the upper Campanian of Alberta and implications for dinosaur biogeography". Cretaceous Research, 51: 292. doi:10.1016/j.cretres.2014.06.011
  32. ^ a b DeCourten and Russell, D. (1985). "A specimen of Ornithomimus velox (Theropoda, Ornithomimidae) from the terminal Cretaceous Kaiparowits Formation of southern Utah." Journal of Paleontology, 59(5): 1091-1099.
  33. ^ Lillegraven and Eberle (1999). "Vertebrate faunal changes through Lancian and Puercan time in southern Wyoming." Journal of Paleontology, 73(4): 691-710.
  34. ^ Zanno, L.E., Weirsma, J.P., Loewen, M.A., Sampson, S.D. and Getty, M.A. (2010). A preliminary report on the theropod dinosaur fauna of the late Campanian Kaiparowits Formation, Grand Staircase-Escalante National Monument, Utah." Learning from the Land Symposium: Geology and Paleontology. Washington, DC: Bureau of Land Management.
  35. ^ Xu, L.; Kobayashi, Y.; Lü, J.; Lee, Y. N.; Liu, Y.; Tanaka, K.; Zhang, X.; Jia, S.; Zhang, J. (2011). "A new ornithomimid dinosaur with North American affinities from the Late Cretaceous Qiupa Formation in Henan Province of China". Cretaceous Research. 32 (2): 213. doi:10.1016/j.cretres.2010.12.004.
  36. ^ Palmer, D., ed. (1999). The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals. London: Marshall Editions. p. 109. ISBN 978-1-84028-152-1.
  37. ^ Rothschild, B., Tanke, D. H., and Ford, T. L., 2001, Theropod stress fractures and tendon avulsions as a clue to activity: In: Mesozoic Vertebrate Life, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, p. 331-336.

Further reading

  • Claessens, L., Loewen, M. and Lavender, Z. 2011. A reevaluation of the genus Ornithomimus based on new preparation of the holotype of O. velox and new fossil discoveries. Journal of Vertebrate Paleontology, SVP Program and Abstracts Book, 2011, pp. 90.
  • Reisdorf, A.G., and Wuttke, M. 2012. Re-evaluating Moodie's Opisthotonic-Posture Hypothesis in fossil vertebrates. Part I: Reptiles - The taphonomy of the bipedal dinosaurs Compsognathus longipes and Juravenator starki from the Solnhofen Archipelago (Jurassic, Germany). Palaeobiodiversity and Palaeoenvironments, doi:10.1007/s12549-011-0068-y
Alvarezsauridae

Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, a consensus of recent work suggests that they evolved from an early branch of maniraptoran theropods. Alvarezsaurids were highly specialized. They had tiny but stout forelimbs, with compact, bird-like hands. Their skeletons suggest that they had massive breast and arm muscles, possibly adapted for digging or tearing. They had long, tube-shaped snouts filled with tiny teeth. They may have been adapted to prey on colonial insects such as termites.

Alvarezsaurus, the type genus of the family, was named for the historian Gregorio Álvarez

Archaeornithomimus

Archaeornithomimus (meaning "ancient bird mimic") is a genus of ornithomimosaurian theropod dinosaur from the Late Cretaceous of China, 70 million years ago.

Arkansaurus

Arkansaurus (meaning "Arkansas lizard") is an extinct genus of ornithomimosaurian theropod dinosaur. It lived during the Albian and Aptian stages of the Early Cretaceous. The type and only species is Arkansaurus fridayi.

Caenagnathidae

Caenagnathidae is a family of bird-like maniraptoran theropod dinosaurs from the Late Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and close relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathids had specialized beaks, long necks, and short tails, and would have been covered in feathers. The relationships of caenagnathids were long a puzzle. The family was originally named by Raymond Martin Sternberg in 1940 as a family of flightless birds. The discovery of skeletons of the related oviraptorids revealed that they were in fact non-avian theropods, and the discovery of more complete caenagnathid remains revealed that Chirostenotes pergracilis, originally named on the basis of a pair of hands, and "Ornithomimus" elegans, named from a foot, were caenagnathids as well.

Coelosaurus antiquus

"Coelosaurus" antiquus ("antique hollow lizard") is a species of theropod dinosaurs. It was named by Joseph Leidy in 1865 for two tibiae found in the Navesink Formation of New Jersey.

This species was later reclassified as a member of the genus Ornithomimus in 1979 by Donald Baird and John R. Horner as Ornithomimus antiquus, and this was followed by some later researchers. However, others have not followed this classification, and have noted that there is no justification for the classification of the New Jersey specimens in a genus known only from western North America. David Weishampel in 2004 considered "C." antiquus to be indeterminate among ornithomimosaurs, and therefore a nomen dubium.In 1979, Baird and Horner discovered that the name "Coelosaurus" was preoccupied by another dubious taxon (based on a single vertebra), named Coelosaurus by an anonymous author now known to be Richard Owen in 1854.Ornithomimid material known from the Severn Formation of Maryland and the Mooreville Chalk and Blufftown formations of Alabama and Georgia have also been assigned to this species.

Death pose

Dinosaur and bird fossils are frequently found in a characteristic posture consisting of head thrown back, tail extended, and mouth wide open. The cause of this posture—sometimes called a "death pose"—has been a matter of scientific debate. Traditional explanations ranged from strong ligaments in the animal's neck desiccating and contracting to draw the body into the pose, to water currents randomly arranging the remains in the position.Faux and Padian suggested in 2007 that the live animal was suffering opisthotonus during its death throes, and that the pose is not the result of any post-mortem process at all. They also reject the idea of water as responsible for randomly arranging the bodies in a "death pose", as different parts of the body and the limbs can be in different directions, which they found unlikely to be the result of moving water. They also found that the claim that drying out of ligaments would make the position does not seem believable either.

Alicia Cutler and colleagues from Brigham Young University in Provo, Utah, think it is related to water. In 2012, paleontologists Achim G. Reisdorf and Michael Wuttke published a study regarding death poses. According to the conclusions of this study, the so-called "opisthotonic posture" is not the result of a cerebral illness creating muscle spasms, and also not of a rapid burial. Rather, peri-mortem submersion resulted in buoyancy that enabled the Ligamentum elasticum to pull the head and tail back.

Deinocheiridae

Deinocheiridae is a family of ornithomimosaurian dinosaurs, living in Asia from the Albian until the Maastrichtian. The family was originally named by Halszka Osmólska and Roniewicz in 1970, including only the type genus Deinocheirus. In a 2014 study by Yuong-Nam Lee and colleagues and published in the journal Nature, it was found that Deinocheiridae was a valid family. Lee et al. found that based on a new phylogenetic analysis including the recently discovered complete skeletons of Deinocheirus, the type genus, as well as Garudimimus and Beishanlong, could be placed as a successive group, with Beishanlong as the most primitive and Deinocheirus as most derived. The family Garudimimidae, named in 1981 by Rinchen Barsbold, is now a junior synonym of Deinocheiridae as the latter family includes the type genus of the former. The group existed from 115 to 69 million years ago, with Beishanlong living from 115 to 100 mya, Garudimimus living from 98 to 83 mya, and Deinocheirus living from 71 to 69 mya.

Deinodon

Deinodon (Greek for "terrible tooth") is a dubious tyrannosaurid dinosaur genus containing a single species, Deinodon horridus. D. horridus is known only from a set of teeth found in the Late Cretaceous Judith River Formation of Montana and named by paleontologist Joseph Leidy in 1856. These were the first tyrannosaurid remains to be described and had been collected by Ferdinand Vandeveer Hayden. The teeth of Deinodon were slightly heterodont, and the holotype of Aublysodon can probably be assigned to Deinodon.

Dromiceiomimus

Dromiceiomimus is a genus of ornithomimid theropod from the Late Cretaceous (early Maastrichtian) of Alberta, Canada.

Gallimimus

Gallimimus ( GAL-i-MY-məs) is a genus of theropod dinosaur that lived in what is now Mongolia during the Late Cretaceous period, about seventy million years ago (mya). Gallimimus is the largest known ornithomimid; adults were about 6 metres (20 ft) long, 1.9 metres (6 ft 3 in) tall at the hip and weighed about 440 kilograms (970 lb). As evidenced by its relative Ornithomimus, it would have had feathers. The head was small and light with large eyes that faced to the sides. The snout was long compared to other ornithomimids, although it was broader and more rounded at the tip than in other species. Gallimimus was toothless with a keratinous (horny) beak, and had a delicate lower jaw. Many of the vertebrae had openings that indicate they were pneumatic (air-filled). The neck was proportionally long in relation to the trunk. The hands were proportionally the shortest of any ornithomimosaur and each had three digits with curved claws. The forelimbs were weak while the hindlimbs were proportionally long.

Several fossils in various stages of growth were discovered by Polish-Mongolian expeditions in the Gobi Desert of Mongolia during the 1960s; a large skeleton discovered in this region was made the holotype specimen of the new genus and species Gallimimus bullatus in 1972. The generic name means "chicken mimic", referring to the similarities between its neck vertebrae and those of the Galliformes. The specific name is derived from bulla, a gold capsule worn by Roman youth, in reference to a bulbous structure at the base of the skull of Gallimimus. At the time it was named, the fossils of Gallimimus represented the most complete and best preserved ornithomimid material yet discovered, and the genus remains one of the best known members of the group. The family Ornithomimidae is part of the group Ornithomimosauria, the "ostrich dinosaurs". Anserimimus, also from Mongolia, is thought to have been the closest relative of Gallimimus.

As an ornithomimid, Gallimimus would have been a fleet (or cursorial) animal, using its speed to escape predators; its speed has been estimated at 42–56 km/h (29–34 mph). It may have had good vision and intelligence comparable to ratite birds. Gallimimus may have lived in groups, based on the discovery of several specimens preserved in a bone bed. Various theories have been proposed regarding the diet of Gallimimus and other ornithomimids. The highly mobile neck may have helped locate small prey on the ground, but it may also have been an opportunistic omnivore. It has also been suggested that it used small columnar structures in its beak for filter-feeding in water, though these structures may instead have been ridges used for feeding on tough plant material, indicative of a herbivorous diet. Gallimimus is the most commonly found ornithomimosaur in the Nemegt Formation, where it lived alongside its relatives Anserimimus and Deinocheirus. Gallimimus was featured in the movie Jurassic Park, in a scene that was important to the history of special effects, and in shaping the common conception of dinosaurs as bird-like animals.

Horseshoe Canyon Formation

The Horseshoe Canyon Formation is a stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta. It takes its name from Horseshoe Canyon, an area of badlands near Drumheller.

The Horseshoe Canyon Formation is part of the Edmonton Group and is up to 230 metres (750 ft) thick. It is of Late Cretaceous age, Campanian to early Maastrichtian stage (Edmontonian Land-Mammal Age), and is composed of mudstone, sandstone, carbonaceous shales, and coal seams. A variety of depositional environments are represented in the succession, including floodplains, estuarine channels, and coal swamps, which have yielded a diversity of fossil material. Tidally-influenced estuarine point bar deposits are easily recognizable as Inclined Heterolithic Stratification (IHS). Brackish-water trace fossil assemblages occur within these bar deposits and demonstrate periodic incursion of marine waters into the estuaries.

The Horseshoe Canyon Formation crops out extensively in the area around Drumheller, as well as farther north along the Red Deer River near Trochu and along the North Saskatchewan River in Edmonton. It is overlain by the Battle, Whitemud, and Scollard formations. The Drumheller Coal Zone, located in the lower part of the Horseshoe Canyon Formation, was mined for sub-bituminous coal in the Drumheller area from 1911 to 1979, and the Atlas Coal Mine in Drumheller has been preserved as a National Historic Site. In more recent times, the Horseshoe Canyon Formation has become a major target for coalbed methane (CBM) production.

Dinosaurs found in the Horseshoe Canyon Formation include Albertavenator, Albertosaurus, Anchiceratops, Anodontosaurus, Arrhinoceratops, Atrociraptor, Epichirostenotes, Edmontonia, Edmontosaurus, Hypacrosaurus, Ornithomimus, Pachyrhinosaurus, Parksosaurus, Saurolophus, and Struthiomimus. Other finds have included mammals such as Didelphodon coyi, non-dinosaur reptiles, amphibians, fish, marine and terrestrial invertebrates and plant fossils. Reptiles such as turtles and crocodilians are rare in the Horseshoe Canyon Formation, and this was thought to reflect the relatively cool climate which prevailed at the time. A study by Quinney et al. (2013) however, showed that the decline in turtle diversity, which was previously attributed to climate, coincided instead with changes in soil drainage conditions, and was limited by aridity, landscape instability, and migratory barriers.

Kaiparowits Formation

The Kaiparowits Formation is a sedimentary rock formation found in the Kaiparowits Plateau in Grand Staircase-Escalante National Monument, in the southern part of Utah in the western United States. It is over 2800 feet (850 m) thick, and is Campanian in age. This Upper Cretaceous formation was formed from alluvial floodplains of large rivers in coastal southern Laramidia; sandstone beds are the deposit of rivers, and mudstone beds represent floodplain deposits. It is fossiliferous, with most specimens from the lower half of the formation, but exploration is only comparatively recent, with most work being done since 1982. It has been estimated that less than 10% of the Kaiparowits formation has been explored for fossils. Most fieldwork has been conducted by The Natural History Museum of Utah.

List of Prehistoric Park episodes

The following is a list of episodes of Prehistoric Park.

Maniraptora

Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to Ornithomimus velox. It contains the major subgroups Avialae, Deinonychosauria, Oviraptorosauria and Therizinosauria. Ornitholestes and the Alvarezsauroidea are also often included. Together with the next closest sister group, the Ornithomimosauria, Maniraptora comprises the more inclusive clade Maniraptoriformes. Maniraptorans first appear in the fossil record during the Jurassic Period (see Eshanosaurus), and are regarded as surviving today as living birds.

Maniraptoriformes

Maniraptoriformes is a clade of dinosaurs with pennaceous feathers and wings that contains ornithomimosaurs and maniraptors. This group was named by Thomas Holtz, who defined it as "the most recent common ancestor of Ornithomimus and birds, and all descendants of that common ancestor."

Ornithomimidae

Ornithomimidae (meaning "bird-mimics") is a group of theropod dinosaurs which bore a superficial resemblance to modern ostriches. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia (now Asia and North America), though they have also been reported from the Wonthaggi Formation of Australia. The group first appeared in the Early Cretaceous.

Ornithomimosauria

The Ornithomimosauria, ornithomimosaurs ("bird-mimic lizards") or ostrich dinosaurs are theropod dinosaurs which bore a superficial resemblance to modern ostriches. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia (now Asia, Europe and North America), as well as Africa and possibly Australia. The group first appeared in the Early Cretaceous and persisted until the Late Cretaceous. Primitive members of the group include Nqwebasaurus, Pelecanimimus, Shenzhousaurus, Hexing and Deinocheirus, the arms of which reached 2.4 m (8 feet) in length. More advanced species, members of the family Ornithomimidae, include Gallimimus, Struthiomimus, and Ornithomimus. Some paleontologists, like Paul Sereno, consider the enigmatic alvarezsaurids to be close relatives of the ornithomimosaurs and place them together in the superfamily Ornithomimoidea (see classification below).

Struthiomimus

Struthiomimus (meaning "ostrich mimic", from the Greek στρούθειος/stroutheios meaning "of the ostrich" and μῖμος/mimos meaning "mimic" or "imitator") is a genus of ornithomimid dinosaurs from the late Cretaceous of North America. Ornithomimids were long-legged, bipedal, ostrich-like dinosaurs with toothless beaks. The type species, Struthiomimus altus, is one of the more common small dinosaurs found in Dinosaur Provincial Park; its abundance suggests that these animals were herbivores or omnivores rather than pure carnivores.

Timeline of ornithomimosaur research

This timeline of ornithomimosaur research is a chronological listing of events in the history of paleontology focused on the ornithomimosaurs, a group of bird-like theropods popularly known as the ostrich dinosaurs. Although fragmentary, probable, ornithomimosaur fossils had been described as far back as the 1860s, the first ornithomimosaur to be recognized as belonging to a new family distinct from other theropods was Ornithomimus velox, described by Othniel Charles Marsh in 1890. Thus the ornithomimid ornithomimosaurs were one of the first major Mesozoic theropod groups to be recognized in the fossil record. The description of a second ornithomimosaur genus did not happen until nearly 30 years later, when Henry Fairfield Osborn described Struthiomimus in 1917. Later in the 20th century, significant ornithomimosaur discoveries began occurring in Asia. The first was a bonebed of "Ornithomimus" (now Archaeornithomimus) asiaticus found at Iren Debasu. More Asian discoveries took place even later in the 20th century, including the disembodied arms of Deinocheirus mirificus and the new genus Gallimimus bullatus. The formal naming of the Ornithomimosauria itself was performed by Rinchen Barsbold in 1976.Early research into ornithomimosaur evolution was based on comparative anatomy. In 1972, Dale Russell argued that the Jurassic Elaphrosaurus of Africa was an ancestral relative of ornithomimids. The descriptions of Garudimimus and Harpymimus in the 1980s revealed the existence of primitive ornithomimosaurs outside of the Ornithomimidae proper. Subsequent research and discoveries during the 1990s refined science's knowledge of ornithomimosaur evolution. In 1994, Pelecanimimus polyodon was described from Europe, the first known ornithomimosaur from that continent and apparently a very evolutionarily primitive taxon. From the late 1990s into the early 21st century cladistic evidence mounted against Russell's hypothesis that ornithomimosaurs were descended from a close relative of Elaphrosaurus, and favored an ancestry close to Pelecanimimus. Paleontologists found that within the theropod family tree, ornithomimosaurs were primitive coelurosaurs closely related to, but outside of, the maniraptorans.The juxtaposition of apparent evolutionary affinities to carnivorous dinosaurs with the possession of toothless beaks has led to controversy among paleontologists trying to reconstruct the diet of ornithomimosaurs. Osborn hypothesized in 1917 that ornithomimosaurs may have eaten plants, social insects, or aquatic invertebrates. In the 1970s paleontologists Russell, Halszka Osmolska, and her colleagues considered ornithomimosaurs carnivores that may have fed on insects, small vertebrates, or eggs. In the early to mid 1980s, however Russell and Elizabeth Nicholls began advocating a reinterpretation of ornithomimosaurs as herbivores. With the 1999 report of gastroliths in the new genus Sinornithomimus, came further support for reinterpreting ornithomimosaurs as herbivores or filter feeders rather than carnivores. In 2001, Mark Norell reported a comb-like structure in the beak of Gallimimus that may have been used for filter feeding, bringing renewed credibility to one of Osborn's 1917 hypotheses. If this interpretation of the evidence is correct, Gallimimus would be the largest terrestrial filter feeder in history.

Basal ornithomimosaurs
Ornithomimoidea

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