Maiasaura

Maiasaura (from the Greek "μαία" and the feminine form of Latin saurus, meaning "good mother reptile" or "good mother lizard" ) is a large herbivorous hadrosaurid ("duck-billed") dinosaur genus that lived in the area currently covered by the state of Montana in the Upper Cretaceous Period (mid to late Campanian), about 76.7 million years ago.[1]

The first fossils of Maiasaura were discovered in 1978. The genus was named in 1979. The name refers to the find of nests with eggs, embryos and young animals, in a nesting colony. These showed that Maiasaura fed its young while they were in the nest, the first time such evidence was obtained for a dinosaur. Hundreds of bones of Maiasaura have been dug up.

Maiasaura was about 9 metres (30 ft) long. Young animals walked on their hind legs, adults on all fours. Maiasaura was probably closely related to Brachylophosaurus.

Maiasaura
Temporal range: Late Cretaceous, 76.7–70.6 Ma
Maiasaura skeleton
Mounted cast, Brussels Natural History Museum
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Order: Ornithischia
Suborder: Ornithopoda
Family: Hadrosauridae
Subfamily: Saurolophinae
Tribe: Brachylophosaurini
Genus: Maiasaura
Horner & Makela, 1979
Type species
Maiasaura peeblesorum
Horner & Makela, 1979

Description

Maiasaura BW
Life restoration

Maiasaura were large, attaining a maximum known length of about 9 metres (30 ft). They had a flat beak typical of hadrosaurids, and thick noses. They had a small, spiky crest in front of the eyes. This crest may have been used in headbutting contests between males during the breeding season.[2]

Maiasaura were herbivorous. They were capable of walking both on two (bipedal) or four (quadrupedal) legs. Studies of the stress patterns of healed bones show that young juveniles under four years old walked mainly bipedal, switching to a mainly quadrupedal style of walking when they grew larger.[3] They appeared to have no defense against predators, except, perhaps, its heavy muscular tail and their herd behavior. Herds were extremely large and could have comprised as many as 10,000 individuals.[2] Maiasaura lived in an inland habitat.[4]

Discovery

Maiasaura embryo
Reconstructed cast by Jack Horner of a Maiasaura emerging from its egg

A skull of Maiasaura, specimen PU 22405 (now in the collections of the Yale Peabody Museum of Natural History as YPM PU 22405 following the transfer of the Princeton University vertebrate paleontology collections), was discovered by Laurie Trexler in 1979 and described by dinosaur paleontologists Jack Horner and Robert Makela as the holotype of a new species. They named the type species Maiasaura peeblesorum. The generic name refers to the Greek goddess Maia, the "Good Mother"; to emphasise this, they used the feminine form of saurus: saura. The specific name honours the families of John and James Peebles, on whose land the finds were made.[5] The generic name refers to Marion Brandvold's discovery in 1978 of a nest with remains of eggshells and babies too large to be hatchlings. These discoveries led to others, and the area became known as "Egg Mountain", in rocks of the Two Medicine Formation near Choteau in western Montana. This was the first proof of giant dinosaurs raising and feeding their young.[2]

Over 200 specimens, in all age ranges, have been found.[6] The announcement of the discovery of Maiasaura attracted renewed scientific interest to the Two Medicine Formation and many other new kinds of dinosaurs were discovered as a result of the increased attention.[7] Choteau Maiasaura remains are found in higher strata than their Two Medicine River counterparts.[8]

Classification

Maiasaura peeblesorum cast - University of California Museum of Paleontology - Berkeley, CA - DSC04688
Cast of a juvenile skeleton

The following cladogram of hadrosaurid relationships was published in 2013 by Alberto Prieto-Márquez et al.:[9]

Saurolophinae
Brachylophosaurini

Acristavus gagstarsoni

Brachylophosaurus canadensis

Maiasaura peeblesorum

Shantungosaurus giganteus

Edmontosaurus

Edmontosaurus regalis

Edmontosaurus annectens

Saurolophini

Kerberosaurus manakini

Sabinas OTU

Prosaurolophus maximus

Saurolophus

Saurolophus morrisi

Saurolophus osborni

Saurolophus angustirostris

Kritosaurini

Wulagasaurus dongi

Kritosaurus navajovius

Big Bend UTEP OTU

Secernosaurus koerneri

Willinakaqe salitralensis

Gryposaurus

Gryposaurus latidens

Gryposaurus notabilis

Gryposaurus monumentensis

Palaeobiology

Maiasaurusnest
Reconstructed nest

Maiasaura lived in herds and it raised its young in nesting colonies. The nests in the colonies were packed closely together, like those of modern seabirds, with the gap between the nests being around 7 metres (23 ft); less than the length of the adult animal.[10] The nests were made of earth and contained 30 to 40 eggs laid in a circular or spiral pattern. The eggs were about the size of ostrich eggs.[2]

The eggs were incubated by the heat resulting from rotting vegetation placed into the nest by the parents, rather than a parent sitting on the nest. Upon hatching, fossils of baby Maiasaura show that their legs were not fully developed and thus they were incapable of walking. Fossils also show that their teeth were partly worn, which means that the adults brought food to the nest.[2]

Maiasaura Nest Model.001 - Natural History Museum of London
Reconstruction of a nest with eggs

The hatchlings grew from a size of 16 to 58 inches (41 to 147 cm) long in the span of their first year. At this point, or perhaps after another year, the animal left the nest. This high rate of growth may be evidence of warm bloodedness. The hatchlings had different facial proportions from the adults, with larger eyes and a shorter snout.[2] These features are associated with cuteness, and commonly elicit care from parents in animals dependent on their parents for survival during the early stages of life.

Studies led by Holly Woodward, Jack Horner, Freedman Fowler et al. have given insight into the life history of Maiasaura, resulting in what is perhaps the most detailed life history of any dinosaur known, and to which all others can be compared. From a sample of fifty individual Maiasaura tibiae, it was found that Maiasaurs had a mortality rate of about 89.9% in their first year of life. If the animals survived their second year, their mortality rate would drop to 12.7%. The animals would spend their next six years maturing and growing. Sexual maturity was found to occur in their third year, while skeletal maturity was attained at eight years of age. In their eighth year and beyond, the mortality rate for Maiasaura would spike back to around 44.4%. The studies that followed also found that Maiasaurs were primarily bipedal as juveniles, and switched to a more quadrupedal stance as they aged. It was also found that Maiasaura also included rotting wood in its diet, as well that its environment had a long, dry season prone to drought. The results of the study were published in the journal Palaeobiology on September 3, 2015.[11][12]

Palaeoecology

Maiasaur Pano-v1
Illustration of a herd of Maiasaura walking along a creekbed, as found in the semi-arid Two Medicine Formation fossil bed. This region was characterized by volcanic ash layers and conifer, fern and horsetail vegetation.

Maiasaura is a characteristic fossil of the middle portion (lithofacies 4) of the Two Medicine Formation, dated to about 76.4 million years ago.[1] Maiasaura lived alongside the troodontid Troodon and the hypsilophodont Orodromeus, as well as the dromaeosaurid Bambiraptor and the tyrannosaur Daspletosaurus.[1] Another species of hadrosaurids, referable to the genus Hypacrosaurus, coexisted with Maiasaura for some time, as Hypacrosaurus remains have been found lower in the Two Medicine Formation than was earlier known.[13] The discovery of an additional hadrosaurid, Gryposaurus latidens, in the same range as Maiasaura has shown that the border between hypothesized distinct faunas in the upper and middle is less distinct than once thought.[13] There seems to be a major diversification in ornithischian taxa after the appearance of Maiasaura within the Two Medicine Formation.[13] The thorough examination of strata found along the Two Medicine River (which exposes the entire upper half of the Two Medicine Formation) indicates that the apparent diversification was a real event rather than a result of preservational biases.[13] While Maiasaura has historically been associated with the Two Medicine formation ceratopsid Einiosaurus in a single fauna, this is inaccurate, as Maiasaura is known exclusively from older strata.[14]

See also

Footnotes

  1. ^ a b c Horner, J. R., Schmitt, J. G., Jackson, F., & Hanna, R. (2001). Bones and rocks of the Upper Cretaceous Two Medicine-Judith River clastic wedge complex, Montana. In Field trip guidebook, Society of Vertebrate Paleontology 61st Annual Meeting: Mesozoic and Cenozoic Paleontology in the Western Plains and Rocky Mountains. Museum of the Rockies Occasional Paper (Vol. 3, pp. 3-14).
  2. ^ a b c d e f "Maiasaura," Dodson, et al. (1994); pages 116-117.
  3. ^ Cubo, Jorge; Woodward, Holly; Wolff, Ewan; Horner, John R. (2015). "First Reported Cases of Biomechanically Adaptive Bone Modeling in Non-Avian Dinosaurs". PLoS ONE. 10 (7): e0131131. Bibcode:2015PLoSO..1031131C. doi:10.1371/journal.pone.0131131. PMC 4495995. PMID 26153689.
  4. ^ "Judithian Climax," Lehman (2001); page 315.
  5. ^ Horner, J.R.; Makela, R. (1979). "Nest of juveniles provides evidence of family structure among dinosaurs". Nature. 282 (5736): 296–298. Bibcode:1979Natur.282..296H. doi:10.1038/282296a0.
  6. ^ Horner and Gorman (1988).
  7. ^ "Introduction," Trexler (2001); pages 299-300.
  8. ^ "Faunal Turnover, Migration, and Evolution," Trexler (2001); page 304.
  9. ^ Prieto-Márquez, A.; Wagner, J.R. (2013). "A new species of saurolophine hadrosaurid dinosaur from the Late Cretaceous of the Pacific coast of North America". Acta Palaeontologica Polonica. 58 (2): 255–268. doi:10.4202/app.2011.0049.
  10. ^ Palmer (1999); page 148.
  11. ^ "Largest dinosaur population growth study ever shows how Maiasaura lived and died: Decades of research on Montana's state fossil -- the 'good mother lizard' Maiasaura peeblesorum -- has resulted in the most detailed life history of any dinosaur known".
  12. ^ Woodward, Holly N.; Freedman Fowler, Elizabeth A.; Farlow, James O.; Horner, John R. (2015). "Maiasaura, a model organism for extinct vertebrate population biology: A large sample statistical assessment of growth dynamics and survivorship". Paleobiology. 41 (4): 503–527. doi:10.1017/pab.2015.19.
  13. ^ a b c d "Faunal Turnover, Migration, and Evolution," Trexler (2001); page 306.
  14. ^ Sullivan, R. M.; Lucas, S. G. (2006). "The Kirtlandian land-vertebrate "age"–faunal composition, temporal position and biostratigraphic correlation in the nonmarine Upper Cretaceous of western North America". New Mexico Museum of Natural History and Science Bulletin. 35: 7–29.

References

  • Dodson, Peter & Britt, Brooks & Carpenter, Kenneth & Forster, Catherine A. & Gillette, David D. & Norell, Mark A. & Olshevsky, George & Parrish, J. Michael & Weishampel, David B. The Age of Dinosaurs. Publications International, LTD. p. 116-117. ISBN 0-7853-0443-6.
  • Horner, Jack and Gorman, James. (1988). Digging Dinosaurs: The Search that Unraveled the Mystery of Baby Dinosaurs, Workman Publishing Co.
  • Lehman, T. M., 2001, Late Cretaceous dinosaur provinciality: In: Mesozoic Vertebrate Life, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, pp. 310–328.
  • Palmer, D., ed. (1999). The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals. London: Marshall Editions. p. 148. ISBN 1-84028-152-9.
  • Trexler, D., 2001, Two Medicine Formation, Montana: geology and fauna: In: Mesozoic Vertebrate Life, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, pp. 298–309.
Acristavus

Acristavus is a genus of saurolophine dinosaur. Fossils have been found from the Campanian Two Medicine Formation in Montana and Wahweap Formation in Utah, United States. The type species A. gagslarsoni was named in 2011. Unlike nearly all hadrosaurids except Edmontosaurus, Acristavus lacked ornamentation on its skull. The discovery of Acristavus is paleontologically significant because it supports the position that the ancestor of all hadrosaurids did not possess cranial ornamentation, and that ornamentation was an adaptation that later arose interdependently in the subfamilies Saurolophinae and Lambeosaurinae. It is closely related to Brachylophosaurus and Maiasaura, and was assigned to a new clade called Brachylophosaurini.

Aralosaurus

Aralosaurus (meaning "Aral Sea lizard", because it was found in the Aral Sea - Greek sauros = lizard) was a genus of hadrosaurid dinosaur which lived during the Late Cretaceous Bostobe Formation of what is now Kazakhstan. Aralosaurus was characterized by a small, bony peak on its nose, much like its relatives Maiasaura and Gryposaurus. It was described by Soviet paleontologist A. Rozhdestvensky in 1968.Aralosaurus was a herbivore that lived in the late Cretaceous period, sometime between 93.5 and 85.8 million years ago. Several relatives, such as Jaxartosaurus have also been found in the surrounding area where Aralosaurus was found.

Barsboldia

Barsboldia (meaning "of Barsbold", a well-known Mongolian paleontologist) was a genus of large hadrosaurid dinosaur from the early Maastrichtian Nemegt Formation of Ömnogöv', Mongolia. It is known from a partial vertebral column, partial pelvis, and some ribs.

Bonapartesaurus

Bonapartesaurus is an extinct genus of herbivorous ornithopod dinosaur belonging to Hadrosauridae, which lived in the area of the modern Argentina during the Campanian and Maastrichtian stages of the Late Cretaceous.

Brachylophosaurini

Brachylophosaurini is a tribe of saurolophine hadrosaurs with known material being from N. America and potentially Asia. It contains at least four taxa; Acristavus (from Montana and Utah), Brachylophosaurus (from Montana and Alberta), Maiasaura (also from Montana), and Probrachylophosaurus (also from Montana). A hadrosaur from the Amur river, Wulagasaurus, might be a member of this tribe, but this is disputed. The group was defined by Terry A. Gates and colleagues in 2011.The clade Brachylophosaurini was defined as "Hadrosaurine ornithopods more closely related to Brachylophosaurus, Maiasaura, or Acristavus than to Gryposaurus or Saurolophus".

Campanian

The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous series. Campanian spans the time from 83.6 (± 0.7) to 72.1 (± 0.6) million years ago. It is preceded by the Santonian and it is followed by the Maastrichtian.The Campanian was an age when a worldwide sea level rise covered many coastal areas. The morphology of some of these areas has been preserved: it is an unconformity beneath a cover of marine sedimentary rocks.

Dinosaur Planet (TV series)

Dinosaur Planet is a four-part American nature documentary that aired on the Discovery Channel as a special-two night event on December 14 and 16, 2003. It is hosted by paleontologist Scott Sampson and narrated by actor Christian Slater. It was released on DVD as a two-disc pack on February 17, 2004, and was also released on VHS around the same time.

The format is similar to Discovery's earlier series When Dinosaurs Roamed America. Each episode tells a fictionalized account of a dinosaur from the Late Cretaceous period. The animals are recreated with computer-generated imagery and composited into present-day filmed locations that approximate prehistoric Earth. Periodic interludes (three in each episode) feature Scott Sampson explaining the scientific findings behind the story, also similar to When Dinosaurs Roamed America, but has improved in quality.

Dinosaur Train

Dinosaur Train is an American/Canadian/Singaporean children's animated series created by Craig Bartlett, who also created Hey Arnold! and Ready Jet Go!. The series features a curious young Tyrannosaurus rex named Buddy who, together with his adopted Pteranodon family, takes the Dinosaur Train to explore his time period, and have adventures with all kinds of dinosaurs. It is produced by The Jim Henson Company in association with Media Development Authority, Sparky Animation, FableVision, and Snee-Oosh, Inc. As of 2018, PBS Kids had ordered 11 more episodes, taking the total number of episodes to 100.

Jack Horner (paleontologist)

John Robert Horner (born June 15, 1946) is an American paleontologist most famous for discovering and naming Maiasaura, providing the first clear evidence that some dinosaurs cared for their young. In addition to his paleontological discoveries, Horner served as the technical advisor for all the Jurassic Park films, had a cameo appearance in Jurassic World, and served as a partial inspiration for one of the lead characters of the franchise, Dr. Alan Grant. Horner studied at the University of Montana, although he did not complete his degree due to undiagnosed dyslexia, and was awarded a Doctorate in Science honoris causa. He retired from Montana State University on July 1, 2016 although he claims to have been pushed out of the Museum of the Rockies after having married an undergraduate student and now teaches as a Presidential Fellow at Chapman University.

Paleontology in Montana

Paleontology in Montana refers to paleontological research occurring within or conducted by people from the U.S. state of Montana. The fossil record in Montana stretches all the way back to the Precambrian. During the Late Precambrian, western Montana was covered by a warm, shallow sea where local bacteria formed stromatolites and bottom-dwelling marine life left tracks on the sediment that would later fossilize. This sea remained in place during the early Paleozoic, although withdrew during the Silurian and Early Devonian, leaving a gap in the local rock record until its return. This sea was home to creatures including brachiopods, conodonts, crinoids, fish, and trilobites. During the Carboniferous the state was home to an unusual cartilaginous fish fauna. Later in the Paleozoic the sea began to withdraw, but with a brief return during the Permian.

During the Triassic Montana was again covered in a sea whose inhabitants are the state's only known fossils from this time. Much of Montana remained covered by seawater into the Late Jurassic, but exposed areas were greened by a flora of conifers, cycads, ferns, and ginkgoes. This coastal plain was home to dinosaurs including the ornithopod Camptosaurus, the sauropods Apatosaurus and Diplodocus, and the theropod Allosaurus. During the early Cretaceous the state was home to its first flowering plants as well as predators such as Deinonychus. Later the Western Interior Seaway, home to creatures such as mosasaurs and plesiosaurs, came to cover the state. On land, the duckbilled Maiasaura formed vast nesting colonies. By the end of the Cretaceous Montana was home to some of the most famous dinosaurs; creatures such as Edmontosaurus, Triceratops, and Tyrannosaurus.

During the early Cenozoic the sea began to withdraw from the state. Plants, insects, dogs, and titanotheres are preserved from this time. Later inhabitants would include arctocyonids, insectivores, multituberculates, pantodonts, primates, and taeniodonts. As the Cenozoic proceeded the climate cooled until the Ice Age started and glaciers entered the state. At this time mammoths, musk oxen, and dire wolves lived in the areas of the state not covered by glaciers. Local Native Americans have been devising mythical explanations for fossils or applying them for practical purposes thousands of years. The first scientific collection of local fossils began in the mid 19th century. Other important discoveries include the plant and insect fossils of Ruby Valley and Deinonychus, which triggered the Dinosaur Renaissance. The Cretaceous Duck-billed dinosaur Maiasaura peeblesorum is the Montana state fossil.

Paleontology in the United States

Paleontology in the United States refers to paleontological research occurring within or conducted by people from the United States. Paleontologists have found that at the start of the Paleozoic era, what is now "North" America was actually in the southern hemisphere. Marine life flourished in the country's many seas. Later the seas were largely replaced by swamps, home to amphibians and early reptiles. When the continents had assembled into Pangaea drier conditions prevailed. The evolutionary precursors to mammals dominated the country until a mass extinction event ended their reign.

The Mesozoic era followed and the dinosaurs began their rise to dominance, spreading into the country before Pangaea split up. During the latter Jurassic Morrison Formation dinosaurs lived in the western states. During the Cretaceous, the Gulf of Mexico expanded until it split North America in half. Plesiosaurs and mosasaurs swam in its waters. Later it began to withdraw and the western states were home to the Hell Creek dinosaurs. Another mass extinction ended the reign of the dinosaurs.

The Cenozoic era began afterward. The inland sea of the Cretaceous vanished and mammals came to dominate the land. The western states were home to primitive camels and horses as well as the carnivorous creodonts. Soon mammals had entered the oceans and the early whale Basilosaurus swam the coastal waters of the southeast. Rhino-like titanotheres dominated Oligocene South Dakota. From this point on the climate in the United States cooled until the Pleistocene, when glaciers spread. Saber-toothed cats, woolly mammoths, mastodons, and dire wolves roamed the land. Humans arrived across a land bridge between Siberia and Alaska and may have played a role in hunting these animals into extinction.

Native Americans have been familiar with fossils for thousands of years, but the first major fossil discovery to attract the attention of formally trained scientists were the Ice Age fossils of Kentucky's Big Bone Lick. These fossils were studied by eminent intellectuals like France's George Cuvier and local statesmen like Benjamin Franklin, Thomas Jefferson, and George Washington. By the beginning of the 19th century, Dinosaur footprints were discovered near the country's east coast. Later in the century, as more dinosaur fossils were uncovered, eminent paleontologists Edward Drinker Cope and Othniel Charles Marsh were embroiled in a bitter rivalry to collect the most fossils and name the most new species.

Early in the 20th century major finds continued, such as the Ice Age mammals of the La Brea Tar Pits. Mid-to-late twentieth-century discoveries in the United States triggered the Dinosaur Renaissance as the discovery of the bird-like Deinonychus overturned misguided notions of dinosaurs as plodding lizard-like animals, highlighting their sophisticated physiology and apparent relationship with birds. Other notable finds in the United States include Maiasaura, which provided early evidence for parental care in dinosaurs and "Seismosaurus", the largest known dinosaur.

Probrachylophosaurus

Probrachylophosaurus bergei is a species of large herbivorous brachylophosaurin hadrosaurid dinosaur known from the Late Cretaceous Campanian Judith River Formation, of Montana.

The significance of this particular hadrosaur taxon is that it is a transitional species between the genera Acristavus and Brachylophosaurus evolving from a crestless ancestor (the former genus) to its crested descendant (the latter genus) while changing the morphology of its nasal bones.

Prosaurolophus

Prosaurolophus (; meaning "before Saurolophus", in comparison to the later dinosaur with a similar head crest) is a genus of hadrosaurid (or duck-billed) dinosaur from the Late Cretaceous of North America. It is known from the remains of at least 25 individuals belonging to two species, including skulls and skeletons, but it remains obscure. Around 9 m (30 ft), its fossils have been found in the late Campanian-age Upper Cretaceous Dinosaur Park Formation in Alberta, and the roughly contemporaneous Two Medicine Formation in Montana, dating to around 75.5-74.0 million years ago. Its most recognizable feature is a small solid crest formed by the nasal bones, sticking up in front of the eyes.

The type species is P. maximus, described by American paleontologist Barnum Brown of the American Museum of Natural History in 1916. A second species, P. blackfeetensis, was described by Jack Horner of the Museum of the Rockies in 1992. The two species were differentiated mainly by crest size and skull proportions.

Saurolophinae

Saurolophinae is a subfamily of hadrosaurid dinosaurs. It has since the mid-20th century generally been called the Hadrosaurinae, a group of largely non-crested hadrosaurs related to the crested sub-family Lambeosaurinae. However, the name Hadrosaurinae is based on the genus Hadrosaurus which was found in more recent studies to be more primitive than either lambeosaurines or other traditional "hadrosaurines", like Edmontosaurus and Saurolophus. As a result of this, the name Hadrosaurinae was dropped or restricted to Hadrosaurus alone, and the subfamily comprising the traditional "hadrosaurines" was renamed the Saurolophinae. Recent phylogenetic work by Hai Xing indicates that Hadrosaurus is placed within the monophyletic group containing all non-lambeosaurine hadrosaurids. Under this view, the traditional Hadrosaurinae is resurrected, with the Hadrosauridae being divided into two clades: Hadrosaurinae and Lambeosaurinae.

Saurolophinae was first defined as a clade in a 2010 phylogenetic analysis by Prieto-Márquez. Traditionally, the "crestless" branch of the family Hadrosauridae had been named Hadrosaurinae. However, the use of the term Hadrosaurinae was questioned in a comprehensive study of hadrosaurid relationships by Albert Prieto-Márquez in 2010. Prieto-Márquez noted that, though the name Hadrosaurinae had been used for the clade of mostly crestless hadrosaurids by nearly all previous studies, its type species, Hadrosaurus foulkii, has almost always been excluded from the clade that bears its name, in violation of the rules for naming animals set out by the ICZN. Prieto-Márquez (2010) defined Hadrosaurinae as only the lineage containing H. foulkii, and used the name Saurolophinae instead for the traditional grouping.The cladogram below follows Godefroit et al. (2012) analysis.

The following cladogram was recovered in the 2013 phylogenetic analysis by Prieto-Márquez (the relationships within Lambeosaurinae and between basal hadrosauroids aren't shown).

Shantungosaurus

Shantungosaurus, meaning "Shandong Lizard", is a genus of saurolophine hadrosaurid dinosaurs found in the Late Cretaceous Wangshi Group of the Shandong Peninsula in China. The stratigraphic interval of Shantungosaurus ranges from the top of the Xingezhuang Formation to the middle of the Hongtuya Formation, middle to late Campanian in age. Shantungosaurus is so far the largest hadrosauroid taxon in the world: the greatest length of its femur is about 1.7 m, and the greatest length of its humerus is about 0.97 m.

Timeline of hadrosaur research

This timeline of hadrosaur research is a chronological listing of events in the history of paleontology focused on the hadrosauroids, a group of herbivorous ornithopod dinosaurs popularly known as the duck-billed dinosaurs. Scientific research on hadrosaurs began in the 1850s, when Joseph Leidy described the genera Thespesius and Trachodon based on scrappy fossils discovered in the western United States. Just two years later he published a description of the much better-preserved remains of an animal from New Jersey that he named Hadrosaurus.The early 20th century saw such a boom in hadrosaur discoveries and research that paleontologists' knowledge of these dinosaurs "increased by virtually an order of magnitude" according to a 2004 review by Horner, Weishampel, and Forster. This period is known as the great North American Dinosaur rush because of the research and excavation efforts of paleontologists like Brown, Gilmore, Lambe, Parks, and the Sternbergs. Major discoveries included the variety of cranial ornamentation among hadrosaurs as scientist came to characterize uncrested, solid crested, and hollow crested species. Notable new taxa included Saurolophus, Corythosaurus, Edmontosaurus, and Lambeosaurus. In 1942 Richard Swann Lull and Wright published what Horner, Weishampel, and Forster characterized as the "first important synthesis of hadrosaurid anatomy and phylogeny".More recent discoveries include gigantic hadrosaurs like Shantungosaurus giganteus from China. At 15 meters in length and nearly 16 metric tons in weight it is the largest known hadrosaur and is known from a nearly complete skeleton.Hadrosaur research has continued to remain active even into the new millennium. In 2000, Horner and others found that hatchling Maiasaura grew to adult body sizes at a rate more like a mammal's than a reptile. That same year, Case and others reported the discovery of hadrosaur bones in Vega Island, Antarctica. After decades of such dedicated research, hadrosaurs have become one of the best understood group of dinosaurs.

Two Medicine Formation

The Two Medicine Formation is a geologic formation, or rock body, that was deposited between 83.5 ± 0.7 Ma and 70.6 ± 3.4 Ma (million years ago), during Campanian (Late Cretaceous) time, and is located in northwestern Montana and southern Alberta. It crops out to the east of the Rocky Mountain Overthrust Belt, and the western portion (about 600 metres (2,000 ft) thick) of this formation is folded and faulted while the eastern part, which thins out into the Sweetgrass Arch, is mostly undeformed plains. Below the formation are the nearshore (beach and tidal zone) deposits of the Virgelle Sandstone, and above it is the marine Bearpaw Shale. Throughout the Campanian, the Two Medicine Fm. was deposited between the western shoreline of the Late Cretaceous Interior Seaway and the eastward advancing margin of the Cordilleran Overthrust Belt. The Two Medicine Fm. is mostly sandstone, deposited by rivers and deltas.

Wulagasaurus

Wulagasaurus (meaning "Wulaga lizard", in reference to the discovery locality) is a genus of saurolophine hadrosaurid dinosaur from the Late Cretaceous of Heilongjiang, China. Its remains were found in a bonebed in the middle Maastrichtian-age Yuliangze Formation, dated to 69 million years ago. This bonebed is otherwise dominated by fossils of the lambeosaurine hadrosaurid (hollow-crested duckbill) Sahaliyania. Wulagasaurus was named by Pascal Godefroit and colleagues in 2008. Only partial remains are known at this time. It is one of several hadrosaurids from the Amur River region named since 2000. The type and only species to date is W. dongi, named in honor of Chinese paleontologist Dong Zhiming.

Wulagasaurus is based on GMH W184, a partial dentary (toothbearing bone of the lower jaw). Godefroit and colleagues assigned additional remains from the bonebed to their new genus, including three braincases, a cheekbone, two maxillae (the toothbearing bone of the upper jaw), another dentary, two shoulder blades, two sternal elements, two upper arm bones, and an ischium. It can be distinguished from other hadrosaurids by its slender dentary and the unique form of its upper arm, which had distinctive articulations and placements for muscle attachments. Godefroit and colleagues performed a phylogenetic analysis that suggests Wulagasaurus was the most basal saurolophine known (which would result in a long ghost lineage), and interpreted this as evidence that saurolophines and hadrosaurids in general originated in Asia, which has been supported by other finds since. As a hadrosaurid, Wulagasaurus would have been an herbivore.In recent studies conducted by researchers from the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP), along with others from Chinese Academy of Science, American Museum of Natural History, and Geological Museum of Heilongjiang Provinces, re-evaluated and re-described Wulagasaurus dongi. Based on both original and recent specimens, they concluded that Wulagasaurus shared many morphological similarities with North American taxon's Brachylophosaurus and Maiasaura, possibly forming a clade-structure within the already existing clade Brachylophosaurini. This hypothesis has been demonstrated by another phylogenetic analysis recently coming out.

You Are Umasou

You Are Umasou (おまえうまそうだな, Omae Umasō da na) is a Japanese picture book series by Tatsuya Miyanishi, published by Poplar. The series has spawned animated film and anime adaptations.

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