Greenish warbler

The greenish warbler (Phylloscopus trochiloides) is a widespread leaf warbler with a breeding range in northeastern Europe and temperate to subtropical continental Asia. This warbler is strongly migratory and winters in India. It is not uncommon as a spring or early autumn vagrant in Western Europe and is annually seen in Great Britain. In Central Europe large numbers of vagrant birds are encountered in some years; some of these may stay to breed, as a handful of pairs does each year in Germany.[2]

Like all leaf warblers, it was formerly placed in the "Old World warbler" assemblage, but now belongs to the new leaf-warbler family Phylloscopidae.[3] The genus name Phylloscopus is from Ancient Greek phullon, "leaf", and skopos, "seeker" (from skopeo, "to watch"). The specific trochiloides is from Ancient Greek trokhalos, "bowed", and -oides "resembling", from the similarity to the willow warbler, P. trochilus.[4] The English name of this species provides a perfect argument in favour of the capitalisation of species names (i.e. treating them as proper nouns), a convention which is generally applied in scientific literature. The decapitalised "greenish warbler" is equally descriptive of many bird species across multiple families, whereas a capitalised "Greenish Warbler" shows unambiguously that Phylloscopus trochiloides is under discussion.

Greenish warbler
Greenish Warbler Sikkim India 11.05.2014
nominate race P. trochiloides trochiloides adult from Pangolakha Wildlife Sanctuary, Sikkim, India
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Passeriformes
Family: Phylloscopidae
Genus: Phylloscopus
P. trochiloides
Binomial name
Phylloscopus trochiloides
(Sundevall, 1837)

5, and see text

Description and ecology

Greenish Warbler
Greenish warbler P.trochiloides from Anamalai Hills, Southern Western Ghats, India
Phylloscopus trochiloides NAUMANN
Western greenish warbler,
P. (t.) viridianus

This is a typical leaf warbler in appearance, grayish-green above and off-white below. The single wing bar found in the southern and western populations distinguishes them from most similar species (except Arctic warbler P. borealis). It is slightly smaller than that species and has a thinner bill, without a dark tip to the lower mandible. A latitude-based analysis of wintering birds indicated that more northerly P. trochiloides are smaller, i.e. this species does not seem to follow Bergmann's rule.[5]

Its song is a high jerky trill, in some populations containing a sequence of down- and more rarely up slurred notes.

It breeds in lowland deciduous or mixed forest; non-breeding birds in the warmer parts of its range may move to montane habitat in summer. Individuals from southeast of the Himalayas are for example quite often seen in Bhutan during the hot months, typically in humid Bhutan Fir (Abies densa) forest up to about 3,800 meters ASL or more, but they do not breed there and return again to the adjacent subtropical lowlands in winter.[6]

The nest is on the ground in low shrub. Like its relatives, this small passerine is insectivorous.

Subspecies and evolution

Greenish warbler ring
Presumed evolution around Himalayas.
Yellow: P. t. trochiloides
Orange: P. t. obscuratus
Red: P. t. plumbeitarsus
green: P. t. "ludlowi"
Blue: P. t. viridanus
P. t. nitidus of the Caucasus is not shown.

It has a number of subspecies, of which P. t . viridianus is the most familiar in Europe. As it seems, it is a ring species, with populations diverging east- and westwards of the Tibetan Plateau, later meeting on the northern side. Their relationships are therefore fairly confusing:[7]

  • Eastern group: greenish warblers
    • Phylloscopus trochiloides trochiloides: greenish warbler
      • Southern rim of the Himalaya eastwards from Nepal into W China.
      • Dusky greyish green above, often traces of second wing bar.
    • Phylloscopus trochiloides obscuratus: dull-green warbler
  • Western group: green warblers
    • Phylloscopus trochiloides viridanus: western greenish warbler
      • Breeds Western Siberia to north-east Europe; at east of range south to NW India.
      • Dull green above, with yellowish supercilium, throat, breast and faint wing bar.

The groups' origin lies probably in the Himalayan region, where trochiloides is found. This taxon is close to the parapatric obscuratus, and to plumbeitarsus which is geographically separated from obscuratus; they all can (and in the case of the former two do naturally) hybridize. P. t. plumbeitarsus is often split as distinct species, as it does not hybridize with viridianus in the narrow zone in the western Sayan Mountains where their ranges overlap.

But phylogenetically, the western taxa are even more distinct. However, there is some gene flow between trochiloides and viridianus also, with their hybrids being especially common in Baltistan; they were once considered another subspecies ludlowi. The green warbler P. nitidus, now by many considered a distinct species, is a mountain isolate that diverged from ancestral viridianus.

Song structure differs mainly between greenish warbler and two-barred warbler, which was formerly considered conspecific. The former has a fairly uniform, long, and warbling song. Around the Himalayas, song structure is similar, but songs are generally shorter. Two-barred warbler, on the other hand, has a long song that can be clearly divided into a warbling part, followed by series of up- and downslurred notes. The songs of obscuratus and "ludlowi", are short, but contain the downslur elements too; in the latter, they uniquely appear at the start of the song.[8]


  1. ^ BirdLife International (2012). "Phylloscopus trochiloides". IUCN Red List of Threatened Species. Version 2013.2. International Union for Conservation of Nature. Retrieved 26 November 2013.
  2. ^ Snow et al. (1998), Töpfer (2007)
  3. ^ Alström et al. (2006)
  4. ^ Jobling (2010)
  5. ^ Katti & Price (2003)
  6. ^ Inskipp et al. (2000)
  7. ^ Snow et al. (1998), Alström (2006)
  8. ^ Irwin (2000)


External links

Ackerdijkse Plassen

Ackerdijkse Plassen is a nature reserve in Oude Leede, a village in the municipality of Pijnacker-Nootdorp, (south of Delft and west of Berkel en Rodenrijs). It is one of the most important bird areas in the Netherlands. In addition to 115 species of breeding birds, the ponds are visited annually by about eighty species of migratory birds, such as greenish warbler, Greater short-toed lark, short-eared owl, booted eagle, lesser spotted eagle, black crowned night heron and osprey. Konik provide grazing in the area.

The area contains a restored farmhouse from 1660.

The area is closed to visitors, however there are hiking trails and bike paths around the outside of the area.

The Nature Society took over the management of the area from the Birdlife Netherlands.

Alan Dean (ornithologist)

Alan R. Dean is a British ornithologist with a special interest in gulls and warblers. He lives in Solihull, West Midlands.

He is a member of the West Midland Bird Club and was an assistant editor of its Annual Reports for 1975-1980. He was a member of the British Birds Rarities Committee from 1984–1992 and in 2007 published a comprehensive account of the committee's history and publications. He has published a number of identification papers in British Birds. His papers and articles on Siberian chiffchaff have been of particular note.

In 2010, the West Midland Bird Club conferred on him life membership, in recognition of his contribution to ornithology.

Arctic warbler

The Arctic warbler (Phylloscopus borealis) is a widespread leaf warbler in birch or mixed birch forest near water throughout its breeding range in Fennoscandia and northern Asia. It has established a foothold in North America, breeding in Alaska. This warbler is strongly migratory; the entire population winters in southeast Asia. It therefore has one of the longest migrations of any Old World insectivorous bird.

It traditionally included populations that breed in Kamchatka, the Kuril Islands and Japan, but genetic and vocal evidence strongly suggested these should be treated as separate species, and are all now considered distinct with the Kamchatka leaf warbler in Kamchatka, Hokkaido and the Kuril Islands, and the Japanese leaf warbler in Japan (except Hokkaido).The nest is on the ground in a low shrub. Like most Old World warblers, this small passerine is insectivorous.

This is a typical leaf warbler in appearance, greyish-green above and off-white below. Its single wing bar distinguishes it from most similar species except the greenish warbler, Phylloscopus trochiloides. It is larger than that species and has a heavier, dagger-like bill, with a dark tip to the lower mandible. Its song is a fast trill.

This species occurs as an autumn vagrant in western Europe and is annual on Great Britain. There were 225 confirmed arctic warbler sightings in Britain between 1958 and 2001.The genus name Phylloscopus is from Ancient Greek phullon, "leaf", and skopos, "seeker" (from skopeo, "to watch"). The specific borealis is from Latin and means "northern".

Birds of South Asia. The Ripley Guide

Birds of South Asia. The Ripley Guide by Pamela C. Rasmussen and John C. Anderton is a two-volume ornithological handbook, covering the birds of South Asia, published in 2005 (second edition in 2012) by the Smithsonian Institution and Lynx Edicions. The geographical scope of the book covers India, Bangladesh, Pakistan, Sri Lanka, Nepal, Bhutan, Maldives, the Chagos archipelago and Afghanistan (the latter country had been excluded from previous works covering this region). In total, 1508 species are covered (this figure includes 85 hypothetical and 67 'possible' species, which are given only shorter accounts). Two notable aspects of Birds of South Asia are its distribution evidence-base — the book's authors based their distributional information almost completely on museum specimens — and its taxonomic approach, involving a large number of species-level splits.

British Birds Rarities Committee

The British Birds Rarities Committee (BBRC), established in 1959, is the national bird rarities committee for Britain. It assesses claimed sightings of bird species that are rarely seen in Britain, based on descriptions, photographs and video recordings submitted by observers. Its findings are published in an annual report in the journal British Birds.

The BBRC covers around 280 species whose annually recorded sightings in Britain fall below a threshold deemed to signify rarity. Since the establishment of the committee, some previously included species have become more common—or at least better recorded; this has resulted in their removal from the committee's list and their reclassification as "scarce migrants".

The committee has a chairman, a secretary, and ten voting members, and is supported by others who serve in an advisory capacity. Since its inception, a total of 69 people have served on the committee as assessors. In addition to assessing annual records of rare birds, the committee conducts regular reviews of batches of previously accepted records on a species-by-species basis, to ensure that only those consistent with advances in knowledge of bird identification are retained, and to determine the subspecies of accepted records. Several species have been problematic for assessment, and extreme examples have taken more than 20 years from initial observation to acceptance. The committee has been criticised for its approach to assessing records where only one observer was present, for not publishing reasons for rejecting the validity of records, and for placing undue weight on descriptive detail when assessing record submissions. Seabird identification has proved particularly difficult, leading some observers to suggest that the committee sets too high a standard.

Green warbler

The green warbler (Phylloscopus nitidus), also known as green willow warbler or green leaf warbler, is a leaf warbler found in the Caucasus Mountains in southcentral Europe.

Like all leaf warblers, it was formerly placed in the "Old World warbler" assemblage, but now belongs to the new leaf-warbler family Phylloscopidae. The genus name Phylloscopus is from Ancient Greek phullon, "leaf", and skopos, "seeker" (from skopeo, "to watch"). The specific nitidus is from Latin and means "shining".It is most closely related to the greenish warbler but is brighter in colour, and the underside is much more yellow. It has one strong and one faint wing bar, especially in young birds.


Greenishmay be:

Shades of green

Simon Greenish, MBE, a British Chartered Civil Engineer and Director of Bletchley Park


Gugh (; Cornish: Keow, meaning "hedge banks") could be described as the sixth inhabited island of the Isles of Scilly, but is usually included with St Agnes with which it is joined by a sandy tombolo known as "The Bar" when exposed at low tide. The island is only about 1 km (0.62 mi) long and about 0.5 km (0.31 mi) wide, with the highest point, Kittern Hill at 34 m (112 ft). The geology consists of Hercynian granite with shallow podzolic soils on the higher ground and deeper sandy soils on the lower ground. The former Gugh farm is just north of the neck across the middle of the island between the two hills. The two houses were designed and built in the 1920s by Charles Hamlet Cooper. The name is often mispronounced as "Goo", "Guff" or even "Gogh".

The island lies within the Isles of Scilly Heritage Coast, is in the Isles of Scilly Area of Outstanding Natural Beauty and is managed by the Isles of Scilly Wildlife Trust. Vegetation cover is mainly wind-pruned heath or dense bracken and bramble with a small area of coastal grassland formed over blown sand which has accumulated near the bar.

In 2013 the Isles of Scilly Seabird Recovery Project was set up by a number of organisations including the RSPB and the Isles of Scilly Wildlife Trust. The five-year project aims to keep the islands of St Agnes and Gugh brown rat (Rattus norvegicus) free, to help breeding sea birds, which lost 25% of their populations between 1983 and 2006. The rats eat eggs and kill the chicks of those birds that nest in burrows or on the ground. Rat removal began in October 2013 by a team of 30 volunteers led by Wildlife Management International Limited (WMIL) of New Zealand, and there has been no signs of rats on St Agnes and Gugh since December 2013. WMIL will return to the islands to do a final check for rats in 2016.


Jomfruland is a small elongated Norwegian island located off the coast of mainland Kragerø in the county of Telemark, about 2 km south of the island of Stråholmen. It measures about 7.5 km by 1 km. Jomfruland provides shelter to the many islands of the Kragerø archipelago from the Skagerrak seas.

Approximately on the island's center there are two white lighthouses – one old and one new. Only the newer is in use today. The lighthouses are most characteristic and can be seen from all sides. These towers are often referred to as the characterizing feature of Kragerø and the archipelago.

Access to Jomfruland is by water taxi, car ferry, or by private vessel. The island has several guest harbours. The island has several attractions, among which are splendid beaches, developed docks with restaurant and kiosk facilities, rolling stone beaches on the island's north side, as well as good hiking possibilities for tourists and residents alike.

Karanji Lake

Karanji Lake (Kannada: ಕಾರಂಜಿ ಕೆರೆ) is a lake located in the city of Mysore in the state of Karnataka, India. The lake is surrounded by a nature park consisting of a butterfly park and a walk-through aviary. This aviary is the biggest 'walk-through aviary' in India. There is also a museum, the Regional Museum of Natural History which is located on the banks of this lake. The total area of Karanji lake is 90 hectares. While waterspread area is about 55 hectares, the foreshore area measures about 35 hectares. Karanji lake is owned by the Mysore Zoo Authority. Mysore Zoo gets a revenue of an average of Rs. 50000 per day from ticket sales to enthusiasts who visit this lake.

Leaf warbler

Leaf warblers are small insectivorous passerine birds belonging to the genus Phylloscopus. The genus was introduced by the German zoologist Friedrich Boie in 1826. The name Phylloscopus is from Ancient Greek phullon, "leaf", and skopos, "seeker" (from skopeo, "to watch").Leaf warblers were formerly included in the Old World warbler family but are now considered to belong to the family Phylloscopidae, introduced in 2006. The family originally included the genus Seicercus, but all species have been moved to Phylloscopus in the most recent classification. Leaf warblers are active, constantly moving, often flicking their wings as they glean the foliage for insects along the branches of trees and bushes. They forage at various levels within forests, from the top canopy to the understorey. Most of the species are markedly territorial both in their summer and winter quarters. Most are greenish or brownish above and off-white or yellowish below. Compared to some other "warblers", their songs are very simple. Species breeding in temperate regions are usually strongly migratory.

List of birds of Kyrgyzstan

376 bird species have occurred in the Kyrgyz Republic.


MigrantWatch is a citizen science non-governmental organisation project in India for collection of information about bird migration. The organisation was conceived in July 2007 and is coordinated by the Science Programme of the National Centre for Biological Sciences, in association with Indian Birds journal.The goal of the MigrantWatch programme is to collect information on the arrival, presence and departure of migrant birds that spend the winter in India and to assess any changes that occur in the timing of migration. The MigrantWatch program provides a website where registered members can upload observations of migratory bird species,and access all the sighting records and maps with data plotted.

In the first year, the program targeted nine species of migratory birds:

Northern shoveller Anas clypeata

Marsh harrier, Circus aeruginosus

Wood sandpiper Tringa glareola

Common (or barn) swallow Hirundo rustica

Grey wagtail Motacilla cinerea

Brown shrike Lanius cristatus

Black redstart Phoenicurus ochruros

Greenish warbler Phylloscopus trochiloides

Rosy starling Sturnus roseusSubsequently, the list was increased to 30 migratory species.

Ring species

In biology, a ring species is a connected series of neighbouring populations, each of which can interbreed with closely sited related populations, but for which there exist at least two "end" populations in the series, which are too distantly related to interbreed, though there is a potential gene flow between each "linked" population. Such non-breeding, though genetically connected, "end" populations may co-exist in the same region (sympatry) thus closing a "ring". The German term Rassenkreis, meaning a ring of populations, is also used.

Ring species represent speciation and have been cited as evidence of evolution. They illustrate what happens over time as populations genetically diverge, specifically because they represent, in living populations, what normally happens over time between long-deceased ancestor populations and living populations, in which the intermediates have become extinct. The evolutionary biologist Richard Dawkins remarks that ring species "are only showing us in the spatial dimension something that must always happen in the time dimension".Formally, the issue is that interfertility (ability to interbreed) is not a transitive relation; if A can breed with B, and B can breed with C, it does not mean that A can breed with C, and therefore does not define an equivalence relation. A ring species is a species with a counterexample to the transitivity of interbreeding. However, it is unclear whether any of the examples of ring species cited by scientists actually permit gene flow from end to end, with many being debated and contested.


Sattal or Sat Tal (Hindi for "seven lakes") is an interconnected group of seven freshwater lakes situated in the Lower Himalayan Range near Bhimtal, a town of the Nainital district in Uttarakhand, India. During the British Raj, the area had a tea plantation, one of four in the Kumaon area at that time.The lakes sit at an altitude of 1370 metres below lush orchards in the Mehragaon valley.

Set amongst dense forests of oak and pine trees, Sattal is one of the few unspoiled and unpolluted freshwater biomes in India. These lakes are a paradise for migratory birds. It is home to a few camps being operated mostly by local people catering to tourists looking for outdoor vacations.

Siberian chiffchaff

Siberian chiffchaff (Phylloscopus (collybita) tristis) is a leaf-warbler which is usually considered a subspecies of the common chiffchaff, but may be a species in its own right.


In biology, a species ( (listen)) is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined.

All species (except viruses) are given a two-part name, a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes in zoological nomenclature). For example, Boa constrictor is one of four species of the genus Boa.

While the definitions given above may seem adequate, when looked at more closely they represent problematic species concepts. For example, the boundaries between closely related species become unclear with hybridisation, in a species complex of hundreds of similar microspecies, and in a ring species. Also, among organisms that reproduce only asexually, the concept of a reproductive species breaks down, and each clone is potentially a microspecies. Though none of these are entirely satisfactory definitions, scientists and conservationists need a species definition which allows them to work, regardless of the theoretical difficulties. If species were fixed and clearly distinct from one another, there would be no problem, but evolutionary processes cause species to change continually, and to grade into one another.

Species were seen from the time of Aristotle until the 18th century as fixed categories that could be arranged in a hierarchy, the great chain of being. In the 19th century, biologists grasped that species could evolve given sufficient time. Charles Darwin's 1859 book The Origin of Species explained how species could arise by natural selection. That understanding was greatly extended in the 20th century through genetics and population ecology. Genetic variability arises from mutations and recombination, while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures. Genes can sometimes be exchanged between species by horizontal gene transfer; new species can arise rapidly through hybridisation and polyploidy; and species may become extinct for a variety of reasons. Viruses are a special case, driven by a balance of mutation and selection, and can be treated as quasispecies.

Telkkämäki Nature Reserve

The Telkkämäki Nature Reserve (in Finnish: Telkkämäen luonnonsuojelualue or Telkkämäen kaskiperinnetila) is an open-air museum and a heritage farm in the municipality of Kaavi, in the Northern Savonia region of Finland. It covers one square kilometer (0.39 sq mi).

The reserve was established in 1989, allowing visitors to see how people lived and farmed when slash-and-burn agriculture was practiced in Eastern Finland in the 15th century. Some areas of Telkkämäki are still burned annually.

The area is managed by a state-owned enterprise, Metsähallitus, which manages most of the protected areas in Finland.

Two-barred warbler

The two-barred warbler (Phylloscopus plumbeitarsus) is a species of leaf warbler (family Phylloscopidae). It was formerly included in the "Old World warbler" assemblage. It is closely related to the greenish warbler, to which it was formerly considered conspecific.

It is found in northern Mongolia, Manchuria and southern Siberia.


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