Gorilla

Gorillas are ground-dwelling, predominantly herbivorous apes that inhabit the forests of central Sub-Saharan Africa. The genus Gorilla is divided into two species: the eastern gorillas and the western gorillas (both critically endangered), and either four or five subspecies. They are the largest living primates. The DNA of gorillas is highly similar to that of humans, from 95 to 99% depending on what is included, and they are the next closest living relatives to humans after the chimpanzees and bonobos.

Gorillas' natural habitats cover tropical or subtropical forests in Sub-Saharan Africa. Although their range covers a small percentage of Sub-Saharan Africa, gorillas cover a wide range of elevations. The mountain gorilla inhabits the Albertine Rift montane cloud forests of the Virunga Volcanoes, ranging in altitude from 2,200 to 4,300 metres (7,200 to 14,100 ft). Lowland gorillas live in dense forests and lowland swamps and marshes as low as sea level, with western lowland gorillas living in Central West African countries and eastern lowland gorillas living in the Democratic Republic of the Congo near its border with Rwanda.[2]

Gorillas[1]
Male gorilla in SF zoo
Western gorilla
(Gorilla gorilla)
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Primates
Suborder: Haplorhini
Infraorder: Simiiformes
Family: Hominidae
Subfamily: Homininae
Tribe: Gorillini
Genus: Gorilla
I. Geoffroy Saint-Hilaire, 1852
Type species
Troglodytes gorilla
Savage, 1847
Species

Gorilla gorilla
Gorilla beringei

Distibución gorilla
Distribution of gorillas
Synonyms
  • Pseudogorilla Elliot, 1913

Etymology

The word "gorilla" comes from the history of Hanno the Navigator, (c. 500 BC) a Carthaginian explorer on an expedition on the west African coast to the area that later became Sierra Leone.[3] Members of the expedition encountered "savage people, the greater part of whom were women, whose bodies were hairy, and whom our interpreters called Gorillae".[4] The word was then later used as the species name, though it is unknown whether what these ancient Carthaginians encountered were truly gorillas, another species of ape or monkeys, or humans.[5]

The American physician and missionary Thomas Staughton Savage and naturalist Jeffries Wyman first described the western gorilla (they called it Troglodytes gorilla) in 1847 from specimens obtained in Liberia.[6] The name was derived from Ancient Greek Γόριλλαι (gorillai), meaning 'tribe of hairy women',[7] described by Hanno.

Evolution and classification

The closest relatives of gorillas are the other two Homininae genera, chimpanzees and humans, all of them having diverged from a common ancestor about 7 million years ago.[8] Human gene sequences differ only 1.6% on average from the sequences of corresponding gorilla genes, but there is further difference in how many copies each gene has.[9] Until recently, gorillas were considered to be a single species, with three subspecies: the western lowland gorilla, the eastern lowland gorilla and the mountain gorilla.[5][10] There is now agreement that there are two species, each with two subspecies. More recently, a third subspecies has been claimed to exist in one of the species. The separate species and subspecies developed from a single type of gorilla during the Ice Age, when their forest habitats shrank and became isolated from each other.[2]

Primatologists continue to explore the relationships between various gorilla populations.[5] The species and subspecies listed here are the ones upon which most scientists agree.

Taxonomy of genus Gorilla[1] Phylogeny of superfamily Hominoidea[11](Fig. 4)
 Hominoidea

humans (genus Homo)

chimpanzees (genus Pan)

gorillas (genus Gorilla)

orangutans (genus Pongo)

gibbons (family Hylobatidae)

The proposed third subspecies of Gorilla beringei, which has not yet received a trinomen, is the Bwindi population of the mountain gorilla, sometimes called the Bwindi gorilla.

Some variations that distinguish the classifications of gorilla include varying density, size, hair colour, length, culture, and facial widths.[2] Population genetics of the lowland gorillas suggest that the western and eastern lowland populations diverged ~261 thousand years ago.[12]

Physical characteristics

Gorilla Male perspective 5
Male gorilla skull

Gorillas move around by knuckle-walking, although they sometimes walk bipedally for short distances while carrying food or in defensive situations,[13] and some Mountain Gorillas use other parts of their hand to aid locomotion (studies of 77 Mountain Gorillas published in 2018 showed 61% only used knuckle walking, but the remainder used knuckle walking plus other parts of their hand—fist walking in ways that do not use the knuckles, using the backs of their hand, and using their palms).[14] Wild male gorillas weigh 136 to 195 kg (300 to 430 lb), while adult females usually weigh about half as much as adult males at 68–113 kg (150–250 lb).

Gorilla gorilla & Gorilla beringei
Western gorilla (Gorilla gorilla) and Eastern gorilla (Gorilla beringei)

Adult males are 1.4 to 1.8 m (4 ft 7 in to 5 ft 11 in) tall, with an arm span that stretches from 2.3 to 2.6 m (7 ft 7 in to 8 ft 6 in). Female gorillas are shorter at 1.25 to 1.5 m (4 ft 1 in to 4 ft 11 in), with smaller arm spans.[15][16][17][18][19] Groves (1970) calculates that average weight of the 47 wild adult male gorillas is 143 kg, while Smith and Jungers(1997) found that the average weight of the 19 wild adult male gorillas is 169 kg.[20] Adult male gorillas are known as silverbacks due to the characteristic silver hair on their backs reaching to the hips. The tallest gorilla recorded was a 1.95 m (6 ft 5 in) silverback with an arm span of 2.7 m (8 ft 10 in), a chest of 1.98 m (6 ft 6 in), and a weight of 219 kg (483 lb), shot in Alimbongo, northern Kivu in May 1938.[19] The heaviest gorilla recorded was a 1.83 m (6 ft 0 in) silverback shot in Ambam, Cameroon, which weighed 267 kg (589 lb).[19] Males in captivity are noted to be capable of reaching weights up to 310 kg (683 lb).[19] Gorilla facial structure is described as mandibular prognathism, that is, the mandible protrudes farther out than the maxilla. Adult males also have a prominent sagittal crest.

The eastern gorilla is more darkly coloured than the western gorilla, with the mountain gorilla being the darkest of all. The mountain gorilla also has the thickest hair. The western lowland gorilla can be brown or grayish with a reddish forehead. In addition, gorillas that live in lowland forests are more slender and agile than the more bulky mountain gorillas. The eastern gorilla also has a longer face and broader chest than the western gorilla.[21]

Studies have shown gorilla blood is not reactive to anti-A and anti-B monoclonal antibodies, which would, in humans, indicate type O blood. Due to novel sequences, though, it is different enough to not conform with the human ABO blood group system, into which the other great apes fit.[22] Like humans, gorillas have individual fingerprints.[23][24] Their eye colour is dark brown, framed by a black ring around the iris.

Distribution and habitat

Nshongi Gorilla Group-7, by Justin Norton
Young gorilla climbing

Gorillas have a patchy distribution. The range of the two species is separated by the Congo River and its tributaries. The western gorilla lives in west central Africa, while the eastern gorilla lives in east central Africa. Between the species, and even within the species, gorillas live in a variety of habitats and elevations. Gorilla habitat ranges from montane forests to swamps. Eastern gorillas inhabit montane and submontane forests between 650 and 4,000 m (2,130 and 13,120 ft) above sea level.[25] Mountain gorillas live in the montane forests at the higher ends of the elevation range, while eastern lowland gorillas live in submontane forests at the lower ends of the elevation range. In addition, eastern lowland gorillas live in montane bamboo forests, as well as lowland forests ranging from 600–3,308 m (1,969–10,853 ft) in elevation.[26] Western gorillas live in both lowland swamp forests and montane forests, and elevations ranging from sea level to 1,600 m (5,200 ft).[25] Western lowland gorillas live in swamp and lowland forests ranging up to 1,600 m (5,200 ft), and Cross River gorillas live in low-lying and submontane forests ranging from 150–1,600 m (490–5,250 ft).

Nesting

Gorilla nest
Gorilla night nest constructed in a tree

Gorillas construct nests for daytime and night use. Nests tend to be simple aggregations of branches and leaves about 2 to 5 ft (0.61 to 1.52 m) in diameter and are constructed by individuals. Gorillas, unlike chimpanzees or orangutans, tend to sleep in nests on the ground. The young nest with their mothers, but construct nests after three years of age, initially close to those of their mothers.[27] Gorilla nests are distributed arbitrarily and use of tree species for site and construction appears to be opportunistic.[28] Nest-building by great apes is now considered to be not just animal architecture, but as an important instance of tool use.[28]

Food and foraging

A gorilla's day is divided between rest periods and travel or feeding periods. Diets differ between and within species. Mountain gorillas mostly eat foliage, such as leaves, stems, pith, and shoots, while fruit makes up a very small part of their diets.[29] Mountain gorilla food is widely distributed and neither individuals nor groups have to compete with one another. Their home ranges average 3–15 km2 (1.16–5.79 mi2), and their movements range around 500 m (0.31 mi) or less on an average day.[29] Despite eating a few species in each habitat, mountain gorillas have flexible diets and can live in a variety of habitats.[29]

Gorillas-moving
Gorillas moving in habitat
Gorilla gorilla08
Gorilla foraging

Eastern lowland gorillas have more diverse diets, which vary seasonally. Leaves and pith are commonly eaten, but fruits can make up as much as 25% of their diets. Since fruit is less available, lowland gorillas must travel farther each day, and their home ranges vary from 2.7–6.5 km2 (1.04 to 2.51 mi2), with day ranges 154–2,280 m (0.096–1.417 mi). Eastern lowland gorillas will also eat insects, preferably ants.[30] Western lowland gorillas depend on fruits more than the others and they are more dispersed across their range.[31] They travel even farther than the other gorilla subspecies, at 1,105 m (0.687 mi) per day on average, and have larger home ranges of 7–14 km2 (2.70–5.41 mi2).[31] Western lowland gorillas have less access to terrestrial herbs, although they can access aquatic herbs in some areas. Termites and ants are also eaten.

Gorillas rarely drink water "because they consume succulent vegetation that is comprised of almost half water as well as morning dew",[32] although both mountain and lowland gorillas have been observed drinking.

Behaviour

Social structure

Mountain gorilla family

Gorillas live in groups called troops. Troops tend to be made of one adult male or silverback, multiple adult females and their offspring.[33][34][35] However, multiple-male troops also exist.[34] A silverback is typically more than 12 years of age, and is named for the distinctive patch of silver hair on his back, which comes with maturity. Silverbacks also have large canine teeth that also come with maturity. Both males and females tend to emigrate from their natal groups. For mountain gorillas, females disperse from their natal troops more than males.[33][36] Mountain gorillas and western lowland gorillas also commonly transfer to second new groups.[33]

Mature males also tend to leave their groups and establish their own troops by attracting emigrating females. However, male mountain gorillas sometimes stay in their natal troops and become subordinate to the silverback. If the silverback dies, these males may be able to become dominant or mate with the females. This behaviour has not been observed in eastern lowland gorillas. In a single male group, when the silverback dies, the females and their offspring disperse and find a new troop.[36][37] Without a silverback to protect them, the infants will likely fall victim to infanticide. Joining a new group is likely to be a tactic against this.[36][38] However, while gorilla troops usually disband after the silverback dies, female eastern lowlands gorillas and their offspring have been recorded staying together until a new silverback transfers into the group. This likely serves as protection from leopards.[37]

Gorilla gorilla11
Silverback gorilla

The silverback is the center of the troop's attention, making all the decisions, mediating conflicts, determining the movements of the group, leading the others to feeding sites, and taking responsibility for the safety and well-being of the troop. Younger males subordinate to the silverback, known as blackbacks, may serve as backup protection. Blackbacks are aged between 8 and 12 years[35] and lack the silver back hair. The bond that a silverback has with his females forms the core of gorilla social life. Bonds between them are maintained by grooming and staying close together.[39] Females form strong relationships with males to gain mating opportunities and protection from predators and infanticidal outside males.[40] However, aggressive behaviours between males and females do occur, but rarely lead to serious injury. Relationships between females may vary. Maternally related females in a troop tend to be friendly towards each other and associate closely. Otherwise, females have few friendly encounters and commonly act aggressively towards each other.[33]

Females may fight for social access to males and a male may intervene.[39] Male gorillas have weak social bonds, particularly in multiple-male groups with apparent dominance hierarchies and strong competition for mates. Males in all-male groups, though, tend to have friendly interactions and socialise through play, grooming, and staying together,[35] and occasionally they even engage in homosexual interactions.[41] Severe aggression is rare in stable groups, but when two mountain gorilla groups meet, the two silverbacks can sometimes engage in a fight to the death, using their canines to cause deep, gaping injuries.[42]

Competition

One possible predator of gorillas is the leopard. Gorilla remains have been found in leopard scat, but this may be the result of scavenging.[43] When the group is attacked by humans, leopards, or other gorillas, an individual silverback will protect the group, even at the cost of his own life.[44]

Reproduction and parenting

Gorillas in Uganda-3, by Fiver Löcker
Young gorilla riding on mother

Females mature at 10–12 years (earlier in captivity), and males at 11–13 years. A female’s first ovulatory cycle occurs when she is six years of age, and is followed by a two-year period of adolescent infertility.[45] The estrous cycle lasts 30–33 days, with outward ovulation signs subtle compared to those of chimpanzees. The gestation period lasts 8.5 months. Female mountain gorillas first give birth at 10 years of age and have four-year interbirth intervals.[45] Males can be fertile before reaching adulthood. Gorillas mate year round.[46]

Females will purse their lips and slowly approach a male while making eye contact. This serves to urge the male to mount her. If the male does not respond, then she will try to attract his attention by reaching towards him or slapping the ground.[47] In multiple-male groups, solicitation indicates female preference, but females can be forced to mate with multiple males.[47] Males incite copulation by approaching a female and displaying at her or touching her and giving a "train grunt".[46] Recently, gorillas have been observed engaging in face-to-face sex, a trait once considered unique to humans and bonobos.[48]

Moka with baby gorilla at Pittsburgh Zoo 8, 2012-02-17
Mother gorilla with 10-day-old infant

Gorilla infants are vulnerable and dependent, thus mothers, their primary caregivers, are important to their survival.[38] Male gorillas are not active in caring for the young, but they do play a role in socialising them to other youngsters.[49] The silverback has a largely supportive relationship with the infants in his troop and shields them from aggression within the group.[49] Infants remain in contact with their mothers for the first five months and mothers stay near the silverback for protection.[49] Infants suckle at least once per hour and sleep with their mothers in the same nest.[50]

Infants begin to break contact with their mothers after five months, but only for a brief period each time. By 12 months old, infants move up to five meters (16 feet) from their mothers. At around 18–21 months, the distance between mother and offspring increases and they regularly spend time away from each other.[51] In addition, nursing decreases to once every two hours.[50] Infants spend only half of their time with their mothers by 30 months. They enter their juvenile period at their third year, and this lasts until their sixth year. At this time, gorillas are weaned and they sleep in a separate nest from their mothers.[49] After their offspring are weaned, females begin to ovulate and soon become pregnant again.[49][50] The presence of play partners, including the silverback, minimizes conflicts in weaning between mother and offspring.[51]

Communication

Twenty-five distinct vocalisations are recognised, many of which are used primarily for group communication within dense vegetation. Sounds classified as grunts and barks are heard most frequently while traveling, and indicate the whereabouts of individual group members.[52] They may also be used during social interactions when discipline is required. Screams and roars signal alarm or warning, and are produced most often by silverbacks. Deep, rumbling belches suggest contentment and are heard frequently during feeding and resting periods. They are the most common form of intragroup communication.[42]

For this reason, conflicts are most often resolved by displays and other threat behaviours that are intended to intimidate without becoming physical. The ritualized charge display is unique to gorillas. The entire sequence has nine steps: (1) progressively quickening hooting, (2) symbolic feeding, (3) rising bipedally, (4) throwing vegetation, (5) chest-beating with cupped hands, (6) one leg kick, (7) sideways running, two-legged to four-legged, (8) slapping and tearing vegetation, and (9) thumping the ground with palms to end display.[53]

Lifespan

A gorilla's lifespan is normally between 35 and 40 years, although zoo gorillas may live for 50 years or more. Colo, a female western gorilla at the Columbus Zoo and Aquarium was the oldest known gorilla, at 60 years of age when she died on January 17, 2017.[54]

Intelligence

Gorrila tool use-Efi
A female gorilla exhibiting tool use by using a tree trunk as a support whilst fishing herbs

Gorillas are considered highly intelligent. A few individuals in captivity, such as Koko, have been taught a subset of sign language. Like the other great apes, gorillas can laugh, grieve, have "rich emotional lives", develop strong family bonds, make and use tools, and think about the past and future.[55] Some researchers believe gorillas have spiritual feelings or religious sentiments.[2] They have been shown to have cultures in different areas revolving around different methods of food preparation, and will show individual colour preferences.[2]

Tool use

The following observations were made by a team led by Thomas Breuer of the Wildlife Conservation Society in September 2005. Gorillas are now known to use tools in the wild. A female gorilla in the Nouabalé-Ndoki National Park in the Republic of Congo was recorded using a stick as if to gauge the depth of water whilst crossing a swamp. A second female was seen using a tree stump as a bridge and also as a support whilst fishing in the swamp. This means all of the great apes are now known to use tools.[56]

In September 2005, a two-and-a-half-year-old gorilla in the Republic of Congo was discovered using rocks to smash open palm nuts inside a game sanctuary.[57] While this was the first such observation for a gorilla, over 40 years previously, chimpanzees had been seen using tools in the wild 'fishing' for termites. Great apes are endowed with semiprecision grips, and have been able to use both simple tools and even weapons, by improvising a club from a convenient fallen branch, for example.

Scientific study

American physician and missionary Thomas Staughton Savage obtained the first specimens (the skull and other bones) during his time in Liberia.[6] The first scientific description of gorillas dates back to an article by Savage and the naturalist Jeffries Wyman in 1847 in Proceedings of the Boston Society of Natural History,[58][59] where Troglodytes gorilla is described, now known as the western gorilla. Other species of gorilla were described in the next few years.[5]

French explorer Paul du Chaillu at close quarters with a gorilla
Drawing of French explorer Paul Du Chaillu at close quarters with a gorilla

The explorer Paul Du Chaillu was the first westerner to see a live gorilla during his travel through western equatorial Africa from 1856 to 1859. He brought dead specimens to the UK in 1861.[60][61][62]

The first systematic study was not conducted until the 1920s, when Carl Akeley of the American Museum of Natural History traveled to Africa to hunt for an animal to be shot and stuffed. On his first trip, he was accompanied by his friends Mary Bradley, a mystery writer, her husband, and their young daughter Alice, who would later write science fiction under the pseudonym James Tiptree Jr. After their trip, Mary Bradley wrote On the Gorilla Trail. She later became an advocate for the conservation of gorillas, and wrote several more books (mainly for children). In the late 1920s and early 1930s, Robert Yerkes and his wife Ava helped further the study of gorillas when they sent Harold Bigham to Africa. Yerkes also wrote a book in 1929 about the great apes.

After World War II, George Schaller was one of the first researchers to go into the field and study primates. In 1959, he conducted a systematic study of the mountain gorilla in the wild and published his work. Years later, at the behest of Louis Leakey and the National Geographic, Dian Fossey conducted a much longer and more comprehensive study of the mountain gorilla. When she published her work, many misconceptions and myths about gorillas were finally disproved, including the myth that gorillas are violent.

Western lowland gorillas (G. g. gorilla) are believed to be one of the zoonotic origins of HIV/AIDS. The SIVgor Simian immunodeficiency virus that infects them is similar to a certain strain of HIV-1.[63][64][65][66]

Genome sequencing

The gorilla became the next-to-last great ape genus to have its genome sequenced. The first gorilla genome was generated with short read and Sanger sequencing using DNA from a female western lowland gorilla named Kamilah. This gave scientists further insight into the evolution and origin of humans. Despite the chimpanzees being the closest extant relatives of humans, 15% of the human genome was found to be more like that of the gorilla.[67] In addition, 30% of the gorilla genome "is closer to human or chimpanzee than the latter are to each other; this is rarer around coding genes, indicating pervasive selection throughout great ape evolution, and has functional consequences in gene expression."[68] Analysis of the gorilla genome has cast doubt on the idea that the rapid evolution of hearing genes gave rise to language in humans, as it also occurred in gorillas.[69]

Cultural references

Since coming to the attention of western society in the 1860s,[62] gorillas have been a recurring element of many aspects of popular culture and media. For example, gorillas have featured prominently in monstrous fantasy films such as King Kong. Pulp fiction, such as Tarzan and Conan the Barbarian, has featured gorillas as physical opponents of the titular protagonists.

Conservation status

All species (and sub-species) of gorilla are listed as Critically Endangered on the IUCN Red List.[70] Now, over 100,000 western lowland gorillas are thought to exist in the wild, with 4,000 in zoos; eastern lowland gorillas have a population of under 5,000 in the wild and 24 in zoos. Mountain gorillas are the most severely endangered, with an estimated population of about 880 left in the wild and none in zoos.[2][70] Threats to gorilla survival include habitat destruction and poaching for the bushmeat trade. In 2004, a population of several hundred gorillas in the Odzala National Park, Republic of Congo was essentially wiped out by the Ebola virus.[71] A 2006 study published in Science concluded more than 5,000 gorillas may have died in recent outbreaks of the Ebola virus in central Africa. The researchers indicated in conjunction with commercial hunting of these apes, the virus creates "a recipe for rapid ecological extinction".[72] Conservation efforts include the Great Apes Survival Project, a partnership between the United Nations Environment Programme and the UNESCO, and also an international treaty, the Agreement on the Conservation of Gorillas and Their Habitats, concluded under UNEP-administered Convention on Migratory Species. The Gorilla Agreement is the first legally binding instrument exclusively targeting gorilla conservation; it came into effect on 1 June 2008.

See also

References

  1. ^ a b Groves, C.P. (2005). Wilson, D.E.; Reeder, D.M., eds. Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore: Johns Hopkins University Press. pp. 181–182. ISBN 0-801-88221-4. OCLC 62265494.
  2. ^ a b c d e f Prince-Hughes, Dawn (1987). Songs of the Gorilla Nation. Harmony. p. 66. ISBN 978-1-4000-5058-1.
  3. ^ Müller, C. (1855–61). Geographici Graeci Minores. pp. 1.1–14: text and trans. Ed, J. Blomqvist (1979).
  4. ^ "A Carthaginian Exploration of the West African Coast". Archived from the original on 14 March 2017.
  5. ^ a b c d Groves, Colin (2002). "A history of gorilla taxonomy" (PDF). In Andrea B. Taylor & Michele L. Goldsmith. Gorilla Biology: A Multidisciplinary Perspective. Cambridge University Press. pp. 15–34. Archived from the original (PDF) on 26 March 2009.
  6. ^ a b Conniff, Richard (2009). "Discoveringgorilla". Evolutionary Anthropology: Issues, News, and Reviews. 18 (2): 55–61. doi:10.1002/evan.20203.
  7. ^ Γόριλλαι Archived 6 June 2011 at the Wayback Machine, Henry George Liddell, Robert Scott, A Greek-English Lexicon, on Perseus Digital Library
  8. ^ Glazko GV, Nei M (March 2003). "Estimation of divergence times for major lineages of primate species". Mol. Biol. Evol. 20 (3): 424–34. doi:10.1093/molbev/msg050. PMID 12644563. Archived from the original on 29 August 2009.
  9. ^ Goidts V; Armengol L; Schempp W; et al. (March 2006). "Identification of large-scale human-specific copy number differences by inter-species array comparative genomic hybridization". Hum. Genet. 119 (1–2): 185–98. doi:10.1007/s00439-005-0130-9. PMID 16395594.
  10. ^ Stewart, Kelly J.; Pascale Sicotte; Martha M. Robbins (2001). "Mountain Gorillas of the Virungas". Fathom / Cambridge University Press. Archived from the original on 21 September 2008. Retrieved 11 September 2008.
  11. ^ Israfil, H.; Zehr, S. M.; Mootnick, A. R.; Ruvolo, M.; Steiper, M. E. (2011). "Unresolved molecular phylogenies of gibbons and siamangs (Family: Hylobatidae) based on mitochondrial, Y-linked, and X-linked loci indicate a rapid Miocene radiation or sudden vicariance event" (PDF). Molecular Phylogenetics and Evolution. 58 (3): 447–455. doi:10.1016/j.ympev.2010.11.005. PMC 3046308. PMID 21074627. Archived from the original (PDF) on 10 May 2012.
  12. ^ McManus, Kimberly F; Kelley, Joanna L; Song, Shiya; Veeramah, Krishna R; Woerner, August E; Stevison, Laurie S; Ryder, Oliver A; Ape Genome Project, Great; Kidd, Jeffrey M; Wall, Jeffrey D; Bustamante, Carlos D; Hammer, Michael F (2015). "Inference of Gorilla Demographic and Selective History from Whole-Genome Sequence Data". Molecular Biology and Evolution. 32 (3): 600–12. doi:10.1093/molbev/msu394. PMC 4327160. PMID 25534031.
  13. ^ Prince-Hughes, Dawn (1987). Songs of the Gorilla Nation. Harmony. pp. 82–3. ISBN 978-1-4000-5058-1.
  14. ^ Thompson, Nathan E; Ostrofsky, Kelly R; McFarlin, Shannon C; Robbins, Martha M; Stoinski, Tara S; Almécija, Sergio (2018). "Unexpected terrestrial hand posture diversity in wild mountain gorillas". American Journal of Physical Anthropology. 166 (1): 84–94. doi:10.1002/ajpa.23404. PMID 29344933.
  15. ^ Miller, Patricia (1997). Gorillas. p. 64. ISBN 978-0919879898. Archived from the original on 10 May 2016.
  16. ^ Wood, B. A. (1978). "Relationship between body size and long bone lengths in Pan and Gorilla". American Journal of Physical Anthropology. 50 (1): 23–5. doi:10.1002/ajpa.1330500104. PMID 736111.
  17. ^ Leigh, S. R.; Shea, B. T. (1995). "Ontogeny and the evolution of adult body size dimorphism in apes". American Journal of Primatology. 36: 37–60. doi:10.1002/ajp.1350360104.
  18. ^ Tuttle, Russell H. (1986). Apes of the World: Their Social Behavior, Communication, Mentality and Ecology. William Andrew. ISBN 978-0815511045.
  19. ^ a b c d Wood (1983). The Guinness Book of Animal Facts and Feats. Sterling Publishing. ISBN 978-0-85112-235-9.
  20. ^ Gorilla Biology: A Multidisciplinary Perspective
  21. ^ Rowe N. (1996) Pictorial guide to the living primates. East Hampton, NY: Pogonias Pr.
  22. ^ Gamble, K. C.; Moyse, J. A.; Lovstad, J. N.; Ober, C. B.; Thompson, E. E. (2011). "Blood groups in the Species Survival Plan, European Endangered Species Program, and managed in situ populations of bonobo (Pan paniscus), common chimpanzee (Pan troglodytes), gorilla (Gorilla ssp.), and orangutan (Pongo pygmaeus ssp.)" (PDF). Zoo Biology. 30 (4): 427–444. doi:10.1002/zoo.20348. PMC 4258062. PMID 20853409. Archived from the original (pdf) on 20 January 2012. Retrieved 5 January 2012.
  23. ^ "Santa Barbara Zoo – Western Lowland Gorilla". santabarbarazoo.org. Archived from the original on 28 September 2006. Retrieved 10 October 2006.
  24. ^ Gorrilas Archived 1 June 2010 at the Wayback Machine. Smithsonian National Zoological Park
  25. ^ a b Sarmiento EE. (2003) "Distribution, taxonomy, genetics, ecology, and causal links of gorilla survival: the need to develop practical knowledge for gorilla conservation". In: Taylor AB, Goldsmith ML (eds.). Gorilla biology: a multidisciplinary perspective. Cambridge (England): Cambridge Univ Pr. pp. 432–71.
  26. ^ Ilambu O. (2001) "Ecology of Eastern Lowland gorilla: is there enough scientific knowledge to mitigate conservation threats associated with extreme disturbances in its distribution range?" In: The apes: challenges for the 21st century. Conference proceedings; 10 May 2000–13; Brookfield, IL. Chicago: Chicago Zoo Soc. pp. 307–12.
  27. ^ Miller-Schroeder, Patricia (1997). Gorillas. Weigl Educational Publishers Limited. p. 20. ISBN 978-0-919879-89-8. Archived from the original on 27 June 2014. Retrieved 4 July 2011.
  28. ^ a b Marchant, Linda Frances; Nishida, Toshisada (1996). Great ape societies. Cambridge University Press. pp. 226–227. ISBN 978-0-521-55536-4. Archived from the original on 27 June 2014. Retrieved 4 July 2011.
  29. ^ a b c McNeilage A. (2001) "Diet and habitat use of two mountain gorilla groups in contrasting habitats in the Virungas". In: Robbins MM, Sicotte P, Stewart KJ (eds.). Mountain gorillas: three decades of research at Karisoke. Cambridge (England): Cambridge University Press. pp. 265–92.
  30. ^ Yamagiwa J, Mwanza N, Yumoto T, Maruhashi T (1994). "Seasonal change in the composition of the diet of eastern lowland gorillas". Primates. 35: 1–14. doi:10.1007/BF02381481.
  31. ^ a b Tutin CG. (1996) "Ranging and social structure of lowland gorillas in the Lopé Reserve, Gabon". In: McGrew WC, Marchant LF, Nishida T (eds.). Great ape societies. Cambridge (England): Cambridge Univ Press pp. 58–70.
  32. ^ GORILLAS – Diet & Eating Habits Archived 11 August 2011 at the Wayback Machine. Seaworld.org. Retrieved on 2011-09-27.
  33. ^ a b c d Watts DP. (1996) "Comparative socio-ecology of gorillas". In: McGrew WC, Marchant LF, Nishida T (eds.). Great ape societies. Cambridge (England): Cambridge Univ Press. pp. 16-28.
  34. ^ a b Yamagiwa J, Kahekwa J, Kanyunyi Basabose A (2003). "Intra-specific variation in social organization of gorillas: implications for their social evolution". Primates. 44 (4): 359–69. doi:10.1007/s10329-003-0049-5. PMID 12942370.
  35. ^ a b c Robbins MM. (2001) "Variation in the social system of mountain gorillas: the male perspective". In: Robbins MM, Sicotte P, Stewart KJ, editors. Mountain gorillas: three decades of research at Karisoke. Cambridge (England): Cambridge Univ Pr. pp. 29–58.
  36. ^ a b c Stokes EJ, Parnell RJ, Olejniczak C (2003). "Female dispersal and reproductive success in wild western lowland gorillas (Gorilla gorilla gorilla)". Behav Ecol Sociobiol. 54 (4): 329–39. doi:10.1007/s00265-003-0630-3. JSTOR 25063274.
  37. ^ a b Yamagiwa J, Kahekwa J. (2001) "Dispersal patterns, group structure, and reproductive parameters of eastern lowland gorillas at Kahuzi in the absence of infanticide". In: Robbins MM, Sicotte P, Stewart KJ (eds.). Mountain gorillas: three decades of research at Karisoke. Cambridge (England): Cambridge Univ Press. pp. 89–122.
  38. ^ a b Watts DP. (1989). "Infanticide in mountain gorillas: new cases and a reconsideration of the evidence". Ethology. 81: 1–18. doi:10.1111/j.1439-0310.1989.tb00754.x.
  39. ^ a b Watts DP. (2003) "Gorilla social relationships: a comparative review". In: Taylor AB, Goldsmith ML (eds). Gorilla biology: a multidisciplinary perspective. Cambridge (England): Cambridge Univ Press. pp. 302–27.
  40. ^ Watts DP. (2001) "Social relationships of female mountain gorillas". In: Robbins MM, Sicotte P, Stewart KJ (eds.). Mountain gorillas: three decades of research at Karisoke. Cambridge (England): Cambridge Univ Press. pp. 216–40.
  41. ^ Juichi Yamagiwa (1987). "Intra- and Inter-group Interactions of an All-male Group of Virunga Mountain Gorillas (Gorilla gorilla beringei )". Primates. 28 (1): 1–30. doi:10.1007/BF02382180.
  42. ^ a b Fossey, D. (1983). Gorillas in the mist. Boston: Houghton Mifflin Company. ISBN 0-395-28217-9.
  43. ^ Fay JM, Carroll R, Kerbis Peterhans JC, Harris D (1995). "Leopard attack on and consumption of gorillas in the Central African Republic". Journal of Human Evolution. 29 (1): 93–9. doi:10.1006/jhev.1995.1048. Archived from the original on 3 April 2012.
  44. ^ Wildlife: Mountain Gorilla Archived 17 February 2012 at the Wayback Machine. AWF. Retrieved on 2011-09-27.
  45. ^ a b Czekala N, Robbins MM. (2001) "Assessment of reproduction and stress through hormone analysis in gorillas". In: Robbins MM, Sicotte P, Stewart KJ, editors. Mountain gorillas: three decades of research at Karisoke. Cambridge (England): Cambridge Univ Pr. pp. 317–39.
  46. ^ a b Watts DP (1991). "Mountain gorilla reproduction and sexual behavior". Am J Primatol. 24 (3–4): 211–225. doi:10.1002/ajp.1350240307.
  47. ^ a b Sicotte P. (2001) "Female mate choice in mountain gorillas". In: Robbins MM, Sicotte P, Stewart KJ, editors. Mountain gorillas: three decades of research at Karisoke. Cambridge (England): Cambridge Univ Pr. pp. 59–87.
  48. ^ Tuan C. Nguyen. "Caught in the act! Gorillas mate face to face". MSN. Archived from the original on 15 February 2008.
  49. ^ a b c d e Stewart KJ. (2001) "Social relationships of immature gorillas and silverbacks". In: Robbins MM, Sicotte P, Stewart KJ, editors. Mountain gorillas: three decades of research at Karisoke. Cambridge (England): Cambridge Univ Press. ISBN 0521019869. pp. 183–213.
  50. ^ a b c Stewart KJ (1988). "Suckling and lactational anoestrus in wild gorillas (Gorilla gorilla)". J Reprod Fertil. 83 (2): 627–34. doi:10.1530/jrf.0.0830627. PMID 3411555.
  51. ^ a b Fletcher A. (2001) "Development of infant independence from the mother in wild mountain gorillas". In: Robbins MM, Sicotte P, Stewart KJ, editors. Mountain gorillas: three decades of research at Karisoke. Cambridge (England): Cambridge University Press. ISBN 0521019869. pp. 153–82.
  52. ^ Harcourt, A.H., Stewart, K.J., Hauser, M. (1993). "Functions of wild gorilla 'close' calls. I. Repertoire, context, and interspecific comparison". Behaviour. 124: 89. doi:10.1163/156853993X00524.CS1 maint: Multiple names: authors list (link)
  53. ^ Maple, T.L., & Hoff, M.P. (1982). Gorilla Behavior. New York: Van Nostrand Reinhold Company.CS1 maint: Multiple names: authors list (link)
  54. ^ "Columbus Zoo Announces the Death of Colo, World's Oldest Zoo Gorilla". www.columbuszoo.org. Retrieved 12 February 2018.
  55. ^ "Planet of No Apes? Experts Warn It's Close" Archived 10 July 2012 at WebCite CBS News Online, 2007-09-12. Retrieved 2008-03-22.
  56. ^ Breuer T, Ndoundou-Hockemba M, Fishlock V (2005). "First Observation of Tool Use in Wild Gorillas". PLoS Biol. 3 (11): e380. doi:10.1371/journal.pbio.0030380. PMC 1236726. PMID 16187795.
  57. ^ "A Tough Nut to Crack for Evolution". CBS News. 18 October 2005. Archived from the original on 14 November 2006. Retrieved 18 October 2006.
  58. ^ Savage TS. (1847). Communication describing the external character and habits of a new species of Troglodytes (T. gorilla) Archived 4 May 2016 at the Wayback Machine. Boston Soc Nat Hist, pp. 245–247.
  59. ^ Savage TS, Wyman J. (1847). Notice of the external characters and habits of Troglodytes gorilla, a new species of orang from the Gaboon River, osteology of the same Archived 20 May 2016 at the Wayback Machine. Boston J Nat Hist 5:417–443.
  60. ^ McCook, S. (1996). ""It May Be Truth, but It Is Not Evidence": Paul du Chaillu and the Legitimation of Evidence in the Field Sciences" (PDF). Osiris. 11: 177–197. doi:10.1086/368759. hdl:10214/7693.
  61. ^ A History of Museum Victoria: Melbourne 1865: Gorillas at the Museum Archived 27 February 2008 at the Wayback Machine. Museumvictoria.com.au. Retrieved on 2011-09-27.
  62. ^ a b Quammen, David (4 April 2013). "Book Review: Planet of the Ape - 'Between Man and Beast,' by Monte Reel". The New York Times. Archived from the original on 6 April 2013. Retrieved 6 April 2013.
  63. ^ Van Heuverswyn, Fran; Li, Yingying; Neel, Cecile; Bailes, Elizabeth; Keele, Brandon F.; Liu, Weimin; Loul, Severin; Butel, Christelle; Liegeois, Florian; Bienvenue, Yanga; Ngolle, Eitel Mpoudi; Sharp, Paul M.; Shaw, George M.; Delaporte, Eric; Hahn, Beatrice H.; Peeters, Martine (2006). "Human immunodeficiency viruses: SIV infection in wild gorillas". Nature. 444 (7116): 164. Bibcode:2006Natur.444..164V. doi:10.1038/444164a. PMID 17093443.
  64. ^ Plantier, Jean-Christophe; Leoz, Marie; Dickerson, Jonathan E; De Oliveira, Fabienne; Cordonnier, François; Lemée, VéRonique; Damond, Florence; Robertson, David L; Simon, François (2009). "A new human immunodeficiency virus derived from gorillas". Nature Medicine. 15 (8): 871–72. doi:10.1038/nm.2016. PMID 19648927.
  65. ^ Sharp, P. M.; Bailes, E.; Chaudhuri, R. R.; Rodenburg, C. M.; Santiago, M. O.; Hahn, B. H. (2001). "The origins of acquired immune deficiency syndrome viruses: where and when?". Philosophical Transactions of the Royal Society B: Biological Sciences. 356 (1410): 867–76. doi:10.1098/rstb.2001.0863. PMC 1088480. PMID 11405934.
  66. ^ Takebe, Y; Uenishi, R; Li, X (2008). "Global Molecular Epidemiology of HIV: Understanding the Genesis of AIDS Pandemic". HIV-1: Molecular Biology and Pathogenesis. Advances in Pharmacology. Advances in Pharmacology. 56. pp. 1–25. doi:10.1016/S1054-3589(07)56001-1. ISBN 9780123736017. PMID 18086407.
  67. ^ Kate Kelland (7 March 2012). "Gorilla genome sheds new light on human evolution". Reuters. Archived from the original on 8 March 2012. Retrieved 8 March 2012.
  68. ^ Aylwyn Scally; Dutheil, Julien Y.; Hillier, Ladeana W.; Jordan, Gregory E.; Goodhead, Ian; Herrero, Javier; Hobolth, Asger; Lappalainen, Tuuli; Mailund, Thomas; Marques-Bonet, Tomas; McCarthy, Shane; Montgomery, Stephen H.; Schwalie, Petra C.; Tang, Y. Amy; Ward, Michelle C.; Xue, Yali; Yngvadottir, Bryndis; Alkan, Can; Andersen, Lars N.; Ayub, Qasim; Ball, Edward V.; Beal, Kathryn; Bradley, Brenda J.; Chen, Yuan; Clee, Chris M.; Fitzgerald, Stephen; Graves, Tina A.; Gu, Yong; Heath, Paul; et al. (2012). "Insights into hominid evolution from the gorilla genome sequence". Nature. 483 (7388): 169–175. Bibcode:2012Natur.483..169S. doi:10.1038/nature10842. hdl:10261/80788. PMC 3303130. PMID 22398555.
  69. ^ Smith, Kerri (7 March 2012). "Gorilla joins the genome club". Nature. doi:10.1038/nature.2012.10185. Archived from the original on 9 March 2012.
  70. ^ a b "Four out of six great apes one step away from extinction – IUCN Red List". iucn.org. 4 September 2016. Archived from the original on 7 October 2016. Retrieved 15 September 2016.
  71. ^ "Gorillas infecting each other with Ebola". NewScientist.com. 10 July 2006. Archived from the original on 12 July 2006. Retrieved 10 July 2006.
  72. ^ "Ebola 'kills over 5,000 gorillas'". News.bbc.co.uk. 8 December 2006. Archived from the original on 29 March 2009. Retrieved 9 December 2006.

External links

Dian Fossey

Dian Fossey (; January 16, 1932 – c. December 26, 1985) was an American primatologist and conservationist known for undertaking an extensive study of mountain gorilla groups from 1966 until her 1985 murder. She studied them daily in the mountain forests of Rwanda, initially encouraged to work there by paleoanthropologist Louis Leakey. Gorillas in the Mist, a book published two years before her death, is Fossey's account of her scientific study of the gorillas at Karisoke Research Center and prior career. It was adapted into a 1988 film of the same name.Fossey was one of the foremost primatologists in the world, a member of the so-called "Trimates", a group formed of prominent female scientists originally sent by Leakey to study great apes in their natural environments, along with Jane Goodall who studied chimpanzees, and Birutė Galdikas, who studied orangutans. During her time in Rwanda, she actively supported conservation efforts, strongly opposed poaching and tourism in wildlife habitats, and made more people acknowledge sapient gorillas. Fossey was brutally murdered in her cabin at a remote camp in Rwanda in December 1985. It has been theorized that her murder was linked to her conservation efforts.

Gorilla City

Gorilla City is a city appearing in American comic books published by DC Comics. The city, hidden in the jungles of Africa, is home to a race of super-intelligent gorillas, that gained their powers from a meteorite. The supervillain Gorilla Grodd is also from the city. Gorilla City first appears in The Flash vol. 1 #106, (April 1959) and was created by John Broome and Carmine Infantino.

Gorilla Glass

Gorilla Glass is a brand of chemically strengthened glass developed and manufactured by Corning, now in its sixth generation, designed to be thin, light and damage-resistant. Gorilla Glass is unique to Corning, but close equivalents exist, including AGC Inc. Dragontrail and Schott AG Xensation.The alkali-aluminosilicate sheet glass is used primarily as cover glass for portable electronic devices, including mobile phones, portable media players, portable computer displays, and television screens. It is manufactured in Harrodsburg, Kentucky, US; Asan, Korea; and Taiwan.

The glass gains its surface strength, ability to contain flaws, and crack-resistance by being immersed in a proprietary, hot potassium salt ion-exchange bath.Corning experimented with chemically strengthened glass in 1960, as part of a "Project Muscle" initiative. Within a few years they had developed a "muscled glass" marketed as Chemcor. The product was used until the early 1990s in commercial and industrial applications, including automotive, aviation and pharmaceutical uses, notably in approximately one hundred 1968 Dodge Dart and Plymouth Barracuda racing cars, where minimizing the vehicle's weight was essential. Experimentation was revived in 2005, investigating whether the glass could be made thin enough for use in consumer electronics. It was brought into commercial use when Apple asked Corning for a thin, toughened glass to be used in its new iPhone.As of October 2017, some five billion devices globally contain Gorilla Glass. While dominating its market, Gorilla Glass faces varying competition from rivals such as Dragontrail and synthetic sapphire.

Gorilla Grodd

Gorilla Grodd is a fictional supervillain appearing in comic books published by DC Comics, primarily as an enemy of The Flash. The character was created by John Broome and Carmine Infantino, and first appeared in The Flash #106 (May 1959). He is an evil, super-intelligent gorilla who gained mental powers after being exposed to a strange metorite's radiation.

IGN named him 35th of the Top 100 Comic Book Villains. The character appears in various shows seen on The CW's Arrowverse, voiced by David Sobolov and shown in CGI. In his TV appearances, Grodd is said to have acquired his super-mental powers through exposure to the radiation from the particle accelerator explosion.

Gorilla Monsoon

Robert James Marella (June 4, 1937 – October 6, 1999), better known by his ring name of Gorilla Monsoon, was an American professional wrestler, play-by-play commentator, and booker.

Monsoon is famous for his run as a super-heavyweight main eventer, and later as the voice of the World Wrestling Federation, as commentator and backstage manager during the 1980s and 1990s. He also portrayed the on-screen role of WWF President from 1995 to 1997.

In professional wrestling, the staging area just behind the entrance curtain at an event, a position which Marella established and where he could often be found during WWF shows late in his career, is named the "Gorilla Position" in his honor. Although remembered fondly by many viewers, Monsoon was voted Worst Television Announcer a record six times by readers of the Wrestling Observer Newsletter in the publication's annual awards poll.

Gorilla Zoe

Alonzo Mathis, better known by his stage name Gorilla Zoe () is an American rapper best known for being a member of the rap group Boyz N Da Hood. His solo debut album Welcome to the Zoo came out in 2007. His next two albums, Don't Feed Da Animals and King Kong, were released in 2009 and 2011. In 2015, he released two mix tapes: Recovery and Raised in the Jungle. As of 2018 he is an independent artist.

Hominidae

The Hominidae (), whose members are known as great apes or hominids, are a taxonomic family of primates that includes eight extant species in four genera: Pongo, the Bornean, Sumatran and Tapanuli orangutan; Gorilla, the eastern and western gorilla; Pan, the common chimpanzee and the bonobo; and Homo, which includes modern humans and its extinct relatives (e.g., the Neanderthal), and ancestors, such as Homo erectus.Several revisions in classifying the great apes have caused the use of the term "hominid" to vary over time. Its original meaning referred only to humans (Homo) and their closest extinct relatives. That restrictive meaning has now been largely assumed by the term "hominin", which comprises all members of the human clade after the split from the chimpanzees (Pan). The current, 21st-century meaning of "hominid" includes all the great apes including humans. Usage still varies, however, and some scientists and laypersons still use "hominid" in the original restrictive sense; the scholarly literature generally shows the traditional usage until around the turn of the 21st century.Within the taxon Hominidae, a number of extant and known extinct, that is, fossil, genera are grouped with the humans, chimpanzees, and gorillas in the subfamily Homininae; others with orangutans in the subfamily Ponginae (see classification graphic below). The most recent common ancestor of all Hominidae lived roughly 14 million years ago, when the ancestors of the orangutans speciated from the ancestral line of the other three genera. Those ancestors of the family Hominidae had already speciated from the family Hylobatidae (the gibbons), perhaps 15 million to 20 million years ago.

Homininae

Homininae, also called "African hominids" or "African apes", is a subfamily of Hominidae. It includes two tribes, with their extant as well as extinct species: 1) the Hominini tribe (with the genus Homo including modern humans and numerous extinct species; the subtribe Australopithecina, comprising at least two extinct genera; and the subtribe Panina, represented only by the genus Pan, which includes common chimpanzees and bonobos)―and 2) the Gorillini tribe (gorillas). Alternatively, the genus Pan is sometimes considered to belong to its own third tribe, Panini. Homininae comprises all hominids that arose after orangutans (subfamily Ponginae) split from the line of great apes. The Homininae cladogram has three main branches, which lead to gorillas (through the tribe Gorillini), and to humans and chimpanzees via the tribe Hominini and subtribes Hominina and Panina (see the evolutionary tree below). There are two living species of Panina (chimpanzees and bonobos) and two living species of gorillas, but only one extant human species. Traces of hypothetical Homo species, including Homo floresiensis and Homo denisova, have been found with dates as recent as 40,000 years ago. Organisms in this subfamily are described as hominine or hominines (not to be confused with the terms hominins or hominini).

Killing of Harambe

On May 28, 2016, a three-year-old boy climbed into a gorilla enclosure at the Cincinnati Zoo and Botanical Garden and was grabbed and dragged by Harambe, a 17-year-old Western lowland gorilla. Fearing for the boy's life, a zoo worker shot and killed Harambe. The incident was recorded on video and received broad international coverage and commentary, including controversy over the choice to kill Harambe. A number of primatologists and conservationists wrote later that the zoo had no other choice under the circumstances, and that it highlighted the danger of zoo animals in close proximity to humans and the need for better standards of care.

Koko (gorilla)

Hanabiko "Koko" (July 4, 1971 – June 19, 2018) was a female western lowland gorilla known for having learned a large number of hand signs from a modified version of American Sign Language (ASL). Koko was born at the San Francisco Zoo and lived most of her life in Woodside, California, at The Gorilla Foundation's preserve in the Santa Cruz Mountains. The name "Hanabiko" (花火子), lit. 'fireworks child', is of Japanese origin and is a reference to her date of birth, the Fourth of July. Koko gained public attention upon a report of her having adopted a kitten as a pet and creating a name for him.

Her instructor and caregiver, Francine Patterson, reported that Koko had an active vocabulary of more than 1,000 signs of what Patterson calls "Gorilla Sign Language" (GSL). In contrast to other experiments attempting to teach sign language to non-human primates, Patterson simultaneously exposed Koko to spoken English from an early age. It was reported that Koko understood approximately 2,000 words of spoken English, in addition to the signs. Koko's life and learning process has been described by Patterson and various collaborators in books, peer-reviewed scientific articles, and on a website.As with other great-ape language experiments, the extent to which Koko mastered and demonstrated language through the use of these signs is disputed. It is generally accepted that she did not use syntax or grammar, and that her use of language did not exceed that of a young human child. However, she scored between 70 and 90 on various IQ scales, and some experts, including Mary Lee Jensvold, claim that "Koko...[used] language the same way people do".

Magilla Gorilla

Magilla Gorilla is a fictional gorilla and the star of The Magilla Gorilla Show by Hanna-Barbera that aired from 1964 to 1966.

Mountain gorilla

The mountain gorilla (Gorilla beringei beringei) is one of two subspecies of the eastern gorilla (Gorilla beringei). It is listed as Endangered on the IUCN Red List, as the total population is estimated to comprise 1,004 individuals in two populations as of 2018. One population lives in Uganda's Bwindi Impenetrable National Park, and the other in the Virunga Mountains in three adjacent national parks, namely Uganda's Mgahinga Gorilla National Park, Rwanda's Volcanoes National Park, and Virunga National Park in the Democratic Republic of Congo.

Phoenix Suns

The Phoenix Suns are an American professional basketball team based in Phoenix, Arizona. The Suns compete in the National Basketball Association (NBA), as a member of the league's Western Conference Pacific Division, and are the only team in their division not based in California. The Suns play their home games at the Talking Stick Resort Arena.

The franchise began play in 1968 as an expansion team, and their early years were shrouded in mediocrity, but their fortunes changed in the 1970s, where, after partnering long-term guard Dick Van Arsdale and center Alvan Adams with Paul Westphal, the Suns reached the 1976 NBA Finals, in what is considered to be one of the biggest upsets in NBA history. However, after failing to capture a championship, the Suns would rebuild around Walter Davis for a majority of the 1980s, until the acquisition of Kevin Johnson in 1988.

Under Johnson, and after trading for perennial NBA All-Star Charles Barkley, and combined with the output of Tom Chambers and Dan Majerle, the Suns reached the playoffs for a franchise-record thirteen consecutive appearances and remained a regular title contender throughout the 1990s, and reached the 1993 NBA Finals. However, the team would again fail to win a championship, and entered into another period of mediocrity until the early part of the 2000s.

In 2004, the Suns reacquired Steve Nash, and immediately returned into playoff contention. With Nash alongside Shawn Marion and Amar'e Stoudemire, and under head coach Mike D'Antoni, the Suns became renowned worldwide for their quick, dynamic offense, which led them to tie a franchise record in wins in the 2004–05 season. Two more top two Conference placements followed, but the Suns again failed to attain an NBA championship, and were forced into another rebuild.

The Suns own the NBA's seventh-best all-time winning percentage, and have the second highest winning percentage of any teams to have never won an NBA championship. 10 Hall of Famers have played for Phoenix, while two Suns—Barkley and Nash—have won the NBA Most Valuable Player (MVP) award while playing for the team.

Professional wrestling throws

Professional wrestling throws are the application of professional wrestling techniques that involve lifting the opponent up and throwing or slamming them down. They are sometimes also called "power" maneuvers, as they are meant to emphasize a wrestler's strength. Many of these moves are used as finishers by many wrestlers. Many maneuvers are known by several different names. Professional wrestlers frequently give their "finisher" (signature moves that usually result in a win) new names that reflect their gimmick. Moves are listed under general categories whenever possible.

Trinomen

In zoological nomenclature, a trinomen (plural: trinomina), trinominal name, or ternary name, refers to the name of a subspecies. For example: "Gorilla gorilla gorilla" (Savage, 1847) for the western lowland gorilla (genus Gorilla, species western gorilla). Also, "Bison bison bison" (Linnaeus, 1758) for the plains bison (genus Bison, species American bison).

A trinomen is a name with three parts: generic name, specific name and subspecific name. The first two parts alone form the binomen or species name. All three names are typeset in italics, and only the first letter of the generic name is capitalised. No indicator of rank is included: in zoology, subspecies is the only rank below that of species. For example: "Buteo jamaicensis borealis is one of the subspecies of the red-tailed hawk (Buteo jamaicensis)."

In a taxonomic publication, a name is incomplete without an author citation and publication details. This indicates who published the name, in what publication, and the date of the publication.

For example: "Phalacrocorax carbo novaehollandiae (Stephens, 1826)" denotes a subspecies of the great cormorant (Phalacrocorax carbo) introduced by James Francis Stephens in 1826 under the subspecies name novaehollandiae ("of New Holland").

If the generic and specific name have already been mentioned in the same paragraph, they are often abbreviated to initial letters. For example one might write: "The great cormorant Phalacrocorax carbo has a distinct subspecies in Australasia, the black shag P. c. novaehollandiae".

While binomial nomenclature came into being and immediately gained widespread acceptance in the mid-18th century, it was not until the early 20th century that the current unified standard of trinominal nomenclature was agreed upon. This became the standard mainly because of tireless promotion by Elliott Coues – even though trinomina in the modern usage were pioneered in 1828 by Carl Friedrich Bruch and around 1850 was widely used especially by Hermann Schlegel and John Cassin. As late as the 1930s, the use of trinomina was not fully established in all fields of zoology. Thus, when referring especially European works of the preceding era, the nomenclature used is usually not in accord with contemporary standards.

Wakanda

Wakanda () is a fictional country located in Sub-Saharan Africa created by Marvel Comics. It is home to the superhero Black Panther. Wakanda first appeared in Fantastic Four #52 (July 1966), and was created by Stan Lee and Jack Kirby.Wakanda has appeared in various media adaptations, including the 2016 Marvel Cinematic Universe film Captain America: Civil War as well as the 2018 films Black Panther and Avengers: Infinity War.

Western gorilla

The western gorilla (Gorilla gorilla) is a great ape—the type species as well as the most populous species of the genus Gorilla.

Western lowland gorilla

The western lowland gorilla (Gorilla gorilla gorilla) is one of two subspecies of the western gorilla (Gorilla gorilla) that lives in montane, primary and secondary forests and lowland swamps in central Africa in Angola, Cameroon, Central African Republic, Republic of the Congo, Democratic Republic of the Congo, Equatorial Guinea and Gabon. It is the nominate subspecies of the western gorilla, and smallest of the four gorilla subspecies.

The Western lowland gorilla is the only subspecies kept in zoos with the exception of Amahoro, a female Eastern lowland gorilla at Antwerp Zoo and a few Mountain gorillas kept captive in Democratic Republic of the Congo.

Extant species of family Hominidae (great apes)
Hominidae
Extant ape species
Study of apes
Legal and social status
See also

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