Eumetabola is an unranked category of Neoptera. Two large unities known as the Paurometabola and Eumetabola are probably from the adelphotaxa of the Neoptera after exclusion of the Plecoptera. The monophyly of these unities appears to be weakly justified.
|Apis dorsata on Tribulus terrestris|
Endopterygota (from Ancient Greek endon “inner” + pterón, “wing” + New Latin -ota “having”), also known as Holometabola, is a superorder of insects within the infraclass Neoptera that go through distinctive larval, pupal, and adult stages. They undergo a radical metamorphosis, with the larval and adult stages differing considerably in their structure and behaviour. This is called holometabolism, or complete metamorphism.
The Endopterygota are among the most diverse insect superorders, with over 1 million living species divided between 11 orders, containing insects such as butterflies, flies, fleas, bees, ants, and beetles.They are distinguished from the Exopterygota (or Hemipterodea) by the way in which their wings develop. Endopterygota (meaning literally "internal winged forms") develop wings inside the body and undergo an elaborate metamorphosis involving a pupal stage. Exopterygota ("external winged forms") develop wings on the outside their bodies and do not go through a pupal stage. The latter trait is plesiomorphic, however, and not exclusively found in the exopterygotes, but also in groups such as Odonata (dragonflies and damselflies), which are not Neoptera, but more basal among insects.
The earliest endopterygote fossils date from the Carboniferous.Eusociality
Eusociality (from Greek εὖ eu "good" and social), the highest level of organization of sociality, is defined by the following characteristics: cooperative brood care (including care of offspring from other individuals), overlapping generations within a colony of adults, and a division of labor into reproductive and non-reproductive groups. The division of labor creates specialized behavioral groups within an animal society which are sometimes called castes. Eusociality is distinguished from all other social systems because individuals of at least one caste usually lose the ability to perform at least one behavior characteristic of individuals in another caste.
Eusociality exists in certain insects, crustaceans and mammals. It is mostly observed and studied in the Hymenoptera (ants, bees, and wasps) and in Isoptera (termites). A colony has caste differences: Queens and reproductive males take the roles of the sole reproducers, while soldiers and workers work together to create a living situation favorable for the brood. In addition to Hymenoptera and Isoptera, there are two known eusocial vertebrates among rodents: the naked mole-rat and the Damaraland mole-rat. Some shrimps, such as Synalpheus regalis, are also eusocial. E. O. Wilson and others have claimed that humans have evolved a weak form of eusociality (e.g., with menopause), but these arguments have been disputed.Flea
Flea, the common name for the order Siphonaptera, includes 2,500 species of small flightless insects that survive as external parasites of mammals and birds. Fleas live by consuming blood, or hematophagy, from their hosts. Adult fleas grow to about 3 mm (0.12 in) long, are usually brown, and have bodies that are "flattened" sideways or narrow, enabling them to move through their host's fur or feathers. They lack wings, but have strong claws preventing them from being dislodged, mouthparts adapted for piercing skin and sucking blood, and hind legs extremely well adapted for jumping. They are able to leap a distance of some 50 times their body length, a feat second only to jumps made by another group of insects, the superfamily of froghoppers. Fleas' larvae are worm-like with no limbs; they have chewing mouthparts and feed on organic debris left on their host's skin.
The Siphonaptera are most closely related to the snow scorpionflies, or snow fleas in the UK, formally the Boreidae, placing them within the Endopterygote insect order Mecoptera. Fleas arose in the early Cretaceous, most likely as ectoparasites of mammals, before moving on to other groups, including birds. Each species of flea is more or less a specialist with respect to its host animal species: many species never breed on any other host, though some are less selective. Some families of fleas are exclusive to a single host group; for example, the Malacopsyllidae are found only on armadillos, the Ischnopsyllidae only on bats, and the Chimaeropsyllidae only on elephant shrews.
The oriental rat flea, Xenopsylla cheopis, is a vector of Yersinia pestis, the bacterium which causes bubonic plague. The disease was spread to humans by rodents such as the black rat, which were bitten by infected fleas. Major outbreaks included the Plague of Justinian, c. 540 and the Black Death, c. 1350, both of which killed a sizeable fraction of the world's population.
Fleas appear in human culture in such diverse forms as flea circuses, poems like John Donne's erotic The Flea, works of music such as by Modest Mussorgsky, and a film by Charlie Chaplin.Louse
Louse (plural: lice) is the common name for members of the order Phthiraptera, which contains nearly 5,000 species of wingless insect. Lice are obligate parasites, living externally on warm-blooded hosts which include every species of bird and mammal, except for monotremes, pangolins, and bats. Lice are vectors of diseases such as typhus.
Chewing lice live among the hairs or feathers of their host and feed on skin and debris, while sucking lice pierce the host's skin and feed on blood and other secretions. They usually spend their whole life on a single host, cementing their eggs, called nits, to hairs or feathers. The eggs hatch into nymphs, which moult three times before becoming fully grown, a process that takes about four weeks.
Humans host two species of louse—the head louse and the body louse are subspecies of Pediculus humanus; the pubic louse, Pthirus pubis. The body louse has the smallest genome of any known insect; it has been used as a model organism and has been the subject of much research.
Lice were ubiquitous in human society until at least the Middle Ages. They appear in folktales, songs such as The Kilkenny Louse House, and novels such as James Joyce's Finnegans Wake. They commonly feature in the psychiatric disorder delusional parasitosis. A louse was one of the early subjects of microscopy, appearing in Robert Hooke's 1667 book, Micrographia.Neoptera
Neoptera is a classification group that includes most orders of the winged insects, specifically those that can flex their wings over their abdomens. This is in contrast with the more basal orders of winged insects (the "Palaeoptera" assemblage), which are unable to flex their wings in this way.Paraneoptera
Paraneoptera is a monophyletic superorder of insects which includes four orders, the bark lice, true lice, thrips, and hemipterans, the true bugs. The mouthparts of the Paraneoptera reflect diverse feeding habits. Basal groups are microbial surface feeders, whereas more advanced groups feed on plant or animal fluids.Strepsiptera
The Strepsiptera are an endopterygote order of insects with nine extant families that include about 600 described species. They are endoparasites in other insects, such as bees, wasps, leafhoppers, silverfish, and cockroaches. Females of most species never emerge from the host after entering its body, finally dying inside it. The early-stage larvae do emerge because they must find an unoccupied living host, and the short-lived males must emerge to seek a receptive female in her host.The order is not well known to non-specialists, and the nearest they have to a common name is "stylops". The name of the order translates to "twisted wing"', giving rise to another name used for the order, twisted-wing insects.Thrips
Thrips (order Thysanoptera) are minute (most are 1 mm long or less), slender insects with fringed wings and unique asymmetrical mouthparts. Different thrips species feed mostly on plants by puncturing and sucking up the contents, although a few are predators. Approximately 6,000 species have been described. They fly only weakly and their feathery wings are unsuitable for conventional flight; instead, thrips exploit an unusual mechanism, clap and fling, to create lift using an unsteady circulation pattern with transient vortices near the wings.
Many thrips species are pests of commercially important crops. A few species serve as vectors for over 20 viruses that cause plant disease, especially the Tospoviruses. Some species of thrips are beneficial as pollinators or as predators of other insects or mites. In the right conditions, such as in greenhouses, many species can exponentially increase in population size and form large swarms because of a lack of natural predators coupled with their ability to reproduce asexually, making them an irritation to humans. In addition, Thrips are also known to invade houses and infest objects such as furniture, bedding and computer monitors - in the latter case by forcing their way in between the LCD and its glass covering. Their identification to species by standard morphological characters is often challenging.