Ecological resilience

In ecology, resilience is the capacity of an ecosystem to respond to a perturbation or disturbance by resisting damage and recovering quickly. Such perturbations and disturbances can include stochastic events such as fires, flooding, windstorms, insect population explosions, and human activities such as deforestation, fracking of the ground for oil extraction, pesticide sprayed in soil, and the introduction of exotic plant or animal species. Disturbances of sufficient magnitude or duration can profoundly affect an ecosystem and may force an ecosystem to reach a threshold beyond which a different regime of processes and structures predominates.[2] Human activities that adversely affect ecosystem resilience such as reduction of biodiversity, exploitation of natural resources, pollution, land use, and anthropogenic climate change are increasingly causing regime shifts in ecosystems, often to less desirable and degraded conditions.[2][3] Interdisciplinary discourse on resilience now includes consideration of the interactions of humans and ecosystems via socio-ecological systems, and the need for shift from the maximum sustainable yield paradigm to environmental resource management which aims to build ecological resilience through "resilience analysis, adaptive resource management, and adaptive governance".[4]

Lake and Mulga ecosystems with alternative stable states[1]


The concept of resilience in ecological systems was first introduced by the Canadian ecologist C.S. Holling [5] in order to describe the persistence of natural systems in the face of changes in ecosystem variables due to natural or anthropogenic causes. Resilience has been defined in two ways in ecological literature:

  1. as the time required for an ecosystem to return to an equilibrium or steady-state following a perturbation (which is also defined as stability by some authors). This definition of resilience is used in other fields such as physics and engineering, and hence has been termed ‘engineering resilience’ by Holling.[5][6]
  2. as "the capacity of a system to absorb disturbance and reorganize while undergoing change so as to still retain essentially the same function, structure, identity, and feedbacks".[4]

The second definition has been termed ‘ecological resilience’, and it presumes the existence of multiple stable states or regimes.[6]

Some shallow temperate lakes can exist within either clear water regime, which provides many ecosystem services, or a turbid water regime, which provides reduced ecosystem services and can produce toxic algae blooms. The regime or state is dependent upon lake phosphorus cycles, and either regime can be resilient dependent upon the lake's ecology and management.[1][2]

Mulga woodlands of Australia can exist in a grass-rich regime that supports sheep herding, or a shrub-dominated regime of no value for sheep grazing. Regime shifts are driven by the interaction of fire, herbivory, and variable rainfall. Either state can be resilient dependent upon management.[1][2]


Ecologists Brian Walker, C S Holling and others describe four critical aspects of resilience: latitude, resistance, precariousness, and panarchy.

The first three can apply both to a whole system or the sub-systems that make it up.

  1. Latitude: the maximum amount a system can be changed before losing its ability to recover (before crossing a threshold which, if breached, makes recovery difficult or impossible).
  2. Resistance: the ease or difficulty of changing the system; how “resistant” it is to being changed.
  3. Precariousness: how close the current state of the system is to a limit or “threshold.”.[4]
  4. Panarchy: the degree to which a certain hierarchical level of an ecosystem is influenced by other levels. For example, organisms living in communities that are in isolation from one another may be organized differently from the same type of organism living in a large continuous population, thus the community-level structure is influenced by population-level interactions.

Closely linked to resilience is adaptive capacity, which is the property of an ecosystem that describes change in stability landscapes and resilience.[6] Adaptive capacity in socio-ecological systems refers to the ability of humans to deal with change in their environment by observation, learning and altering their interactions.[2]

Human impacts

Resilience refers to ecosystem's stability and capability of tolerating disturbance and restoring itself.  If the disturbance is of sufficient magnitude or duration, a threshold may be reached where the ecosystem undergoes a regime shift, possibly permanently. Sustainable use of environmental goods and services requires understanding and consideration of the resilience of the ecosystem and its limits. However, the elements which influence ecosystem resilience are complicated. For example, various elements such as the water cycle, fertility, biodiversity, plant diversity and climate, interact fiercely and affect different systems.

There are many areas where human activity impacts upon and is also dependent upon the resilience of terrestrial, aquatic and marine ecosystems. These include agriculture, deforestation, pollution, mining, recreation, overfishing, dumping of waste into the sea and climate change.


Agriculture can be seen as a significant example which the resilience of terrestrial ecosystems should be considered. The organic matter (elements carbon and nitrogen) in soil, which is supposed to be recharged by multiple plants, is the main source of nutrients for crop growth.[7] At the same time, intensive agriculture practices in response to global food demand and shortages involves the removal of weeds and the application of fertilisers to increase food production. However, as a result of agricultural intensification and the application of herbicides to control weeds, fertilisers to accelerate and increase crop growth and pesticides to control insects, plant biodiversity is reduced as is the supply of organic matter to replenish soil nutrients and prevent run-off. This leads to a reduction in soil fertility and productivity.[7] More sustainable agricultural practices would take into account and estimate the resilience of the land and monitor and balance the input and output of organic matter.


The term deforestation has a meaning that covers crossing the threshold of forest's resilience and losing its ability to return its originally stable state. To recover itself, a forest ecosystem needs suitable interactions among climate conditions and bio-actions, and enough area. In addition, generally, the resilience of a forest system allows recovery from a relatively small scale of damage (such as lightning or landslide) of up to 10 per cent of its area.[8] The larger the scale of damage, the more difficult it is for the forest ecosystem to restore and maintain its balance.

Deforestation also decreases biodiversity of both plant and animal life and can lead to an alteration of the climatic conditions of an entire area. Deforestation can also lead to species extinction, which can have a domino effect particularly when keystone species are removed or when a significant number of species is removed and their ecological function is lost.[3][9]

Climate change

Climate resilience is generally defined as the capacity for a socio-ecological system to: (1) absorb stresses and maintain function in the face of external stresses imposed upon it by climate change and (2) adapt, reorganize, and evolve into more desirable configurations that improve the sustainability of the system, leaving it better prepared for future climate change impacts. Increasingly, climate change is threatening human communities around the world in a variety of ways such as rising sea levels, increasingly frequent large storms, tidal surges and flooding damage. One of the main results of climate change is rising sea water temperature which has a serious effect on coral reefs, through thermal-stress related coral bleaching. Between 1997-1998 the most significant worldwide coral bleaching event was recorded which corresponded with the El Nino Southern Oscillation, with significant damage to the coral reefs of the Western Indian Ocean.[10]


It has been estimated by the United Nations Food and Agriculture Organisation that over 70% of the world’s fish stocks are either fully exploited or depleted which means overfishing threatens marine ecosystem resilience and this is mostly by rapid growth of fishing technology.[11] One of the negative effects on marine ecosystems is that over the last half-century the stocks of coastal fish have had a huge reduction as a result of overfishing for its economic benefits.[12] Blue fin tuna is at particular risk of extinction. Depletion of fish stocks results in lowered biodiversity and consequently imbalance in the food chain, and increased vulnerability to disease.

In addition to overfishing, coastal communities are suffering the impacts of growing numbers of large commercial fishing vessels in causing reductions of small local fishing fleets. Many local lowland rivers which are sources of fresh water have become degraded because of the inflows of pollutants and sediments.[13]

Dumping of waste into the sea

Dumping both depends upon ecosystem resilience whilst threatening it. Dumping of sewage and other contaminants into the ocean is often undertaken for the dispersive nature of the oceans and adaptive nature and ability for marine life to process the marine debris and contaminants. However, waste dumping threatens marine ecosystems by poisoning marine life and eutrophication.

Poisoning marine life

According to the International Maritime Organisation oil spills can have serious effects on marine life. The OILPOL Convention recognized that most oil pollution resulted from routine shipboard operations such as the cleaning of cargo tanks.  In the 1950s, the normal practice was simply to wash the tanks out with water and then pump the resulting mixture of oil and water into the sea. OILPOL 54   prohibited the dumping of oily wastes within a certain distance from land and in 'special areas' where the danger to the environment was especially acute. In 1962 the limits were extended by means of an amendment adopted at a conference organized by IMO. Meanwhile, IMO in 1965 set up a Subcommittee on Oil Pollution, under the auspices of its Maritime Safety committee, to address oil pollution issues.[14]

The threat of oil spills to marine life is recognised by those likely to be responsible for the pollution, such as the International Tanker Owners Pollution Federation:

The marine ecosystem is highly complex and natural fluctuations in species composition, abundance and distribution are a basic feature of its normal function. The extent of damage can therefore be difficult to detect against this background variability. Nevertheless, the key to understanding damage and its importance is whether spill effects result in a downturn in breeding success, productivity, diversity and the overall functioning of the system. Spills are not the only pressure on marine habitats; chronic urban and industrial contamination or the exploitation of the resources they provide are also serious threats.[15]

Eutrophication and algal blooms

The Woods Hole Oceanographic Institution calls nutrient pollution the most widespread, chronic environmental problem in the coastal ocean. The discharges of nitrogen, phosphorus, and other nutrients come from agriculture, waste disposal, coastal development, and fossil fuel use. Once nutrient pollution reaches the coastal zone, it stimulates harmful overgrowths of algae, which can have direct toxic effects and ultimately result in low-oxygen conditions. Certain types of algae are toxic. Overgrowths of these algae result in harmful algal blooms, which are more colloquially referred to as "red tides" or "brown tides". Zooplankton eat the toxic algae and begin passing the toxins up the food chain, affecting edibles like clams, and ultimately working their way up to seabirds, marine mammals, and humans. The result can be illness and sometimes death.[16]

Sustainable development

There is increasing awareness that a greater understanding and emphasis of ecosystem resilience is required to reach the goal of sustainable development.[13][17][18] A similar conclusion is drawn by Perman et al. who use resilience to describe one of 6 concepts of sustainability; "A sustainable state is one which satisfies minimum conditions for ecosystem resilience through time".[19] Resilience science has been evolving over the past decade, expanding beyond ecology to reflect systems of thinking in fields such as economics and political science. And, as more and more people move into densely populated cities, using massive amounts of water, energy, and other resources, the need to combine these disciplines to consider the resilience of urban ecosystems and cities is of paramount importance.[20]

Academic perspectives

The interdependence of ecological and social systems has gained renewed recognition since the late 1990s by academics including Berkes and Folke[21] and developed further in 2002 by Folke et al.[1] As the concept of sustainable development has evolved beyond the 3 pillars of sustainable development to place greater political emphasis on economic development. This is a movement which causes wide concern in environmental and social forums and which Clive Hamilton describes as "the growth fetish".[22]

The purpose of ecological resilience that is proposed is ultimately about averting our extinction as Walker cites Holling in his paper: "[..] "resilience is concerned with [measuring] the probabilities of extinction” (1973, p. 20)".[23] Becoming more apparent in academic writing is the significance of the environment and resilience in sustainable development. Folke et al state that the likelihood of sustaining development is raised by "Managing for resilience"[1] whilst Perman et al. propose that safeguarding the environment to "deliver a set of services" should be a "necessary condition for an economy to be sustainable".[19]

The flaw of the free market

The challenge of applying the concept of ecological resilience to the context of sustainable development is that it sits at odds with conventional economic ideology and policy making. Resilience questions the free market model within which global markets operate. Inherent to the successful operation of a free market is specialisation which is required to achieve efficiency and increase productivity. This very act of specialisation weakens resilience by permitting systems to become accustomed to and dependent upon their prevailing conditions. In the event of unanticipated shocks; this dependency reduces the ability of the system to adapt to these changes.[1] Correspondingly; Perman et al. note that; "Some economic activities appear to reduce resilience, so that the level of disturbance to which the ecosystem can be subjected to without parametric change taking place is reduced".[19]

Moving beyond sustainable development

Berkes and Folke table a set of principles to assist with "building resilience and sustainability" which consolidate approaches of adaptive management, local knowledge-based management practices and conditions for institutional learning and self-organisation.[21]

More recently, it has been suggested by Andrea Ross that the concept of sustainable development is no longer adequate in assisting policy development fit for today’s global challenges and objectives. This is because the concept of sustainable development is "based on weak sustainability" which doesn’t take account of the reality of "limits to earth's resilience".[24] Ross draws on the impact of climate change on the global agenda as a fundamental factor in the "shift towards ecological sustainability" as an alternative approach to that of sustainable development.[24]

In environmental policy

Scientific research associated with resilience is beginning to play a role in influencing policy-making and subsequent environmental decision making.

This occurs in a number of ways:

  • Observed resilience within specific ecosystems drives management practice. When resilience is observed to be low, or impact seems to be reaching the threshold, management response can be to alter human behavior to result in less adverse impact to the ecosystem.[13]
  • Ecosystem resilience impacts upon the way that development is permitted/environmental decision making is undertaken, similar to the way that existing ecosystem health impacts upon what development is permitted. For instance, remnant vegetation in the states of Queensland and New South Wales are classified in terms of ecosystem health and abundance. Any impact that development has upon threatened ecosystems must consider the health and resilience of these ecosystems. This is governed by the Threatened Species Conservation Act 1995 in New South Wales [25] and the Vegetation Management Act 1999 in Queensland.[26]
  • International level initiatives aim at improving socio-ecological resilience worldwide through the cooperation and contributions of scientific and other experts. An example of such an initiative is the Millennium Ecosystem Assessment [27] whose objective is "to assess the consequences of ecosystem change for human well-being and the scientific basis for action needed to enhance the conservation and sustainable use of those systems and their contribution to human well-being". Similarly, the United Nations Environment Programme [28] aim is "to provide leadership and encourage partnership in caring for the environment by inspiring, informing, and enabling nations and peoples to improve their quality of life without compromising that of future generations.

Environmental management in legislation

Ecological resilience and the thresholds by which resilience is defined are closely interrelated in the way that they influence environmental policy-making, legislation and subsequently environmental management. The ability of ecosystems to recover from certain levels of environmental impact is not explicitly noted in legislation, however, because of ecosystem resilience, some levels of environmental impact associated with development are made permissible by environmental policy-making and ensuing legislation.

Some examples of the consideration of ecosystem resilience within legislation include:

  • Environmental Planning and Assessment Act 1979 (NSW) [29] – A key goal of the Environmental Assessment procedure is to determine whether proposed development will have a significant impact upon ecosystems.
  • Protection of the Environment (Operations) Act 1997 (NSW) [30] – Pollution control is dependent upon keeping levels of pollutants emitted by industrial and other human activities below levels which would be harmful to the environment and its ecosystems. Environmental protection licenses are administered to maintain the environmental objectives of the POEO Act and breaches of license conditions can attract heavy penalties and in some cases criminal convictions.[31]
  • Threatened Species Conservation Act 1995 (NSW) [32] – This Act seeks to protect threatened species while balancing it with development.

See also


  1. ^ a b c d e f Folke, C., Carpenter,S., Elmqvist, T., Gunderson, L., Holling C.S., Walker, B. (2002). "Resilience and Sustainable Development: Building Adaptive Capacity in a World of Transformations". Ambio. 31 (5): 437–440. doi:10.1639/0044-7447(2002)031[0437:rasdba];2. PMID 12374053.CS1 maint: Multiple names: authors list (link)
  2. ^ a b c d e Folke, C.; Carpenter, S.; Walker, B.; Scheffer, M.; Elmqvist, T.; Gunderson, L.; Holling, C.S. (2004). "Regime Shifts, Resilience, and Biodiversity in Ecosystem Management". Annual Review of Ecology, Evolution, and Systematics. 35: 557–581. doi:10.1146/annurev.ecolsys.35.021103.105711.
  3. ^ a b Peterson, G.; Allen, C.R.; Holling, C.S. (1998). "Ecological Resilience, Biodiversity, and Scale". Ecosystems. 1 (1): 6–18. CiteSeerX doi:10.1007/s100219900002.
  4. ^ a b c Walker, B.; Holling, C. S.; Carpenter, S. R.; Kinzig, A. (2004). "Resilience, adaptability and transformability in social–ecological systems". Ecology and Society. 9 (2): 5. doi:10.5751/ES-00650-090205.
  5. ^ a b Holling, C.S. (1973). "Resilience and stability of ecological systems". Annual Review of Ecology and Systematics. 4: 1–23. doi:10.1146/
  6. ^ a b c Gunderson, L.H. (2000). "Ecological Resilience — In Theory and Application". Annual Review of Ecology and Systematics. 31: 425–439. doi:10.1146/annurev.ecolsys.31.1.425.
  7. ^ a b Tilman, D. (May 1999). "Global environmental impacts of agricultural expansion: The need of sustainable and efficient practices". Proc. Natl. Acad. Sci. U.S.A. 96 (11): 5995–6000. doi:10.1073/pnas.96.11.5995. PMC 34218. PMID 10339530.
  8. ^ Davis R.; Holmgren P. (2 November 2000). "On Definitions of Forest and Forest Change". Forest Resources Assessment Programme, Working Paper 33. Food and Agriculture Organization of the United Nation, Forestry Department.
  9. ^ Naik, A. (29 June 2010). "Deforestation Statistics".
  10. ^ Obura, D.O. (2005). "Resilience and climate change: lessons from local reefs and bleaching in the Western Indian Ocean". Estuarine, Coastal and Shelf Science. 63 (3): 353–372. doi:10.1016/j.ecss.2004.11.010.
  11. ^ YPTE 2010 Overfishing: Environmental Facts Young Peoples Trust for the Environment Viewed September 12, 2010. Overfishing: Environmental Facts Archived 2010-11-30 at the Wayback Machine
  12. ^ "Overfishing". Grinning Planet. 2010.
  13. ^ a b c Gibbs, M.T. (2009). "Resilience: What is it and what does it mean for marine policymakers?". Marine Policy. 33 (2): 322–331. doi:10.1016/j.marpol.2008.08.001.
  14. ^ IMO 2010 "Oil Pollution". International Maritime Organization. Archived from the original on 2009-07-07.
  15. ^ ITOPF 2010 "Effects of Oil Spills". International Tanker Owners Pollution Federation.
  16. ^ "Water Pollution Effects". Grinning Planet. 2010.
  17. ^ Walker, B.; Carpenter, S.; et al. (2002). "Resilience Management in Social-ecological Systems: a Working Hypothesis for a Participatory Approach". Conservation Ecology. 6 (1): 14. doi:10.5751/ES-00356-060114.
  18. ^ Brand, F. (2009). "Critical natural capital revisited: Ecological resilience and sustainable development". Ecological Economics. 68 (3): 605–612. doi:10.1016/j.ecolecon.2008.09.013.
  19. ^ a b c Perman, R, Ma, Y, McGilvray, J and M.Common. (2003). “Natural Resource and Environmental Economics”. Longman. 26, 52, 86.
  20. ^ "Ecological and Urban Resilience". 12 October 2011.
  21. ^ a b Berkes, F. and Folke, C., (ed Colding, J.) (1998). “Linking Social and Ecological Systems: Management practices and social mechanisms for building resilience”. Cambridge University Press: 1, 33, 429, 433.
  22. ^ Hamilton, C. (2010). “Requiem for a Species: Why we Resist the Truth about Climate Change”. Earthscan. 32, 14.
  23. ^ Walker, J. (2007). “The Strange Evolution of Holling’s Resilience or The Resilience of Economics and the Eternal Return of Infinite Growth”. Submission to TfC e-Journal. 8
  24. ^ a b Ross A (2008). "Modern Interpretations of Sustainable Development". Journal of Law and Society. 36 (1): 32. doi:10.1111/j.1467-6478.2009.00455.x.
  25. ^ DECCW 2010 "Threatened Species". Climate change and Water. New South Wales Department of the Environment.
  26. ^ DERM 2010 "Vegetation Management: Legislation and Policy". Queensland Department of the Environment and Resource Management. Archived from the original on 2010-07-14.
  27. ^ "Millennium Ecosystem Assessment".
  28. ^ United Nations Environment Programme. Viewed September 12, 2010 United Nations Environment Programme
  29. ^ "Environmental Planning and Assessment Act 1979". No. 203. NSW.
  30. ^ "Protection of the Environment (Operations) Act 1997". No. 156. NSW.
  31. ^ DECCW 2010 "Environment protection licences". Climate change and Water. New South Wales Department of Environment, Climate Change and Water. Archived from the original on 2011-02-25.
  32. ^ Threatened Species Conservation Act 1995 (NSW) No.101 Threatened Species Conservation Act

Further reading

  • Hulme, M. (2009). “Why we Disagree about Climate Change: Understanding Controversy, Inaction and Opportunity". Cambridge University Press.
  • Lee, M. (2005) “EU Environmental Law: Challenges, Change and Decisions Making”. Hart. 26.
  • Maclean K, Cuthill M, Ross H. (2013). Six attributes of social resilience. Journal of Environmental Planning and Management. (online first)
  • Pearce, D.W. (1993). “Blueprint 3: Measuring Sustainable Development”. Earthscan.
  • Andrew Zolli; Ann Marie Healy (2013). Resilience: Why Things Bounce Back. Simon & Schuster. ISBN 978-1451683813.

External links

  • Resilience Alliance — a research network that focuses on social-ecological resilience Resilience Alliance
  • Stockholme Resilience Centre — an international centre that advances trans disciplinary research for governance of social-ecological systems with a special emphasis on resilience — the ability to deal with change and continue to develop Stockholm Resilience Centre
  • TURaS — a European project mapping urban transitioning towards resilience and sustainability TURaS
  • Microdocs:Resilience — a short documentary on resilience Resilience

Adaptability (Latin: adaptō "fit to, adjust") is a feature of a system or of a process. This word has been put to use as a specialised term in different disciplines and in business operations. Word definitions of adaptability as a specialised term differ little from dictionary definitions. According to Andresen and Gronau adaptability in the field of organizational management can in general be seen as an ability to change something or oneself to fit to occurring changes. In ecology, adaptability has been described as the ability to cope with unexpected disturbances in the environment.

With respect to business and manufacturing systems and processes, adaptability has come to be seen increasingly as an important factor for their efficiency and economic success. In contrast, adaptability and efficiency are held to be in opposition to each other in biological and ecological systems, requiring a trade-off, since both are important factors in the success of such systems. To determine the adaptability of a process or a system, it should be validated concerning some criteria.

Bibliography of ecology

This is a bibliography of ecology.

Biodiversity loss

Loss of biodiversity or biodiversity loss is the extinction of species (plant or animal) worldwide, and also the local reduction or loss of species in a certain habitat.

The latter phenomenon can be temporary or permanent, depending on whether the environmental degradation that leads to the loss is reversible through ecological restoration / ecological resilience or effectively permanent (e.g. through land loss). Global extinction has so far been proven to be irreversible.

Even though permanent global species loss is a more dramatic phenomenon than regional changes in species composition, even minor changes from a healthy stable state can have dramatic influence on the food web and the food chain insofar as reductions in only one species can adversely affect the entire chain (coextinction), leading to an overall reduction in biodiversity, possible alternative stable states of an ecosystem notwithstanding. Ecological effects of biodiversity are usually counteracted by its loss. Reduced biodiversity in particular leads to reduced ecosystem services and eventually poses an immediate danger for food security, also for humankind.

C. S. Holling

Crawford Stanley (Buzz) Holling, (born December 6, 1930) is a Canadian ecologist, and Emeritus Eminent Scholar and Professor in Ecological Sciences at the University of Florida. Holling is one of the conceptual founders of ecological economics.

Cuban cuisine

Cuban cuisine is a blend of Spanish, African, and Caribbean cuisines. Some Cuban recipes share spices and techniques with Spanish and African cooking, with some Caribbean influence in spice and flavor. This results in a blend of the several different cultural influences, A small but noteworthy Chinese influence can also be accounted for, mainly in the Havana area. There is also some Italian influence. During colonial times, Cuba was an important port for trade, and many Spaniards who lived there brought their culinary traditions with them

Ecological collapse

Ecological collapse refers to a situation where an ecosystem suffers a drastic, possibly permanent, reduction in carrying capacity for all organisms, often resulting in mass extinction. Usually, an ecological collapse is precipitated by a disastrous event occurring on a short time scale. Ecological collapse can be considered as a consequence of ecosystem collapse on the biotic elements that depended on the original ecosystem.Ecosystems have the ability to rebound from a disruptive agent. The difference between collapse or a gentle rebound is determined by two factors—the toxicity of the introduced element and the resiliency of the original ecosystem.Through natural selection the planet's species have continuously adapted to change through variation in their biological composition and distribution. Mathematically it can be demonstrated that greater numbers of different biological factors tend to dampen fluctuations in each of the individual factors.Scientists can predict tipping points for ecological collapse. The most frequently used model for predicting food web collapse is called R50, which is a reliable measurement model for food web robustness.

Ecological stability

An ecosystem is said to possess ecological stability (or equilibrium) if it does not experience unexpected large changes in its characteristics across time, or if it is capable of returning to its equilibrium state after a perturbation (a capacity known as resilience). Although the terms community stability and ecological stability are sometimes used interchangeably, community stability refers only to the characteristics of communities. It is possible for an ecosystem or a community to be stable in some of their properties and unstable in others. For example, a vegetation community in response to a drought might conserve biomass but lose biodiversity.The concept of ecological stability emerged in the first half of the 20th century. With the advancement of theoretical ecology in the 1970s, the usage of the term has expanded to a wide variety of scenarios. This overuse of the term has led to controversy over its definition and implementation. In 1997, Grimm and Wissel made an inventory of 167 definitions used in the literature and found 70 different stability concepts. One of the strategies that these two authors proposed to clarify the subject is to replace ecological stability with more specific terms, such as constancy, resilience and persistence. Following this strategy, an ecosystem which oscillates cyclically around a fixed point, such as the one delineated by the predator-prey equations, would be described as persistent and resilient, but not as constant. Some authors, however, see good reason for the abundance of definitions, because they reflect the extensive variety of real and mathematical systems.Stable ecological systems abound in nature, and the scientific literature has documented them to a great extent. Scientific studies mainly describe grassland plant communities and microbial communities. Nevertheless, it is important to mention that not every community or ecosystem in nature is stable. Also, noise plays an important role on biological systems and, in some scenarios, it can fully determine their temporal dynamics.

Herald Shoal

Herald Shoal is a region of high benthic productivity on the Chukchi Sea shelf. It serves as rich foraging habitat for many species of marine mammals and birds.

Massachusetts Green High Performance Computing Center

Massachusetts Green High Performance Computing Center (MGHPCC) is an intercollegiate high-performance computing facility located in Holyoke, Massachusetts, connected to that city's municipal fiber grid and powered by Holyoke Gas and Electric via the Holyoke Canal System and Dam. MGHPCC is a joint venture of Boston University, Harvard, MIT, Northeastern, and the University of Massachusetts system; the facility holds the capacity for hundreds of thousands of cores in clusters provided by its affiliates. For example, as of 2016 one cluster used by UMass contained a network of 14,376 cores, both Intel and AMD, and more than 1.1 petabytes of on-site storage on an FDR Infiniband network. The facility maintains capacity for regular expansion, with key partners investing capability upgrades in the current building and more than 4 acres of additional undeveloped space.

Microbial biogeography

Microbial biogeography is a subset of biogeography, a field that concerns the distribution of organisms across space and time. Although biogeography traditionally focused on plants and larger animals, recent studies have broadened this field to include distribution patterns of microorganisms. This extension of biogeography to smaller scales—known as "microbial biogeography"—is enabled by ongoing advances in genetic technologies.

The aim of microbial biogeography is to reveal where microorganisms live, at what abundance, and why. Microbial biogeography can therefore provide insight into the underlying mechanisms that generate and hinder biodiversity. Microbial biogeography also enables predictions of where certain organisms can survive and how they respond to changing environments, making it applicable to several other fields such as climate change research.

Nina-Marie Lister

Nina-Marie Lister is currently an Associate Professor and the Graduate Program Director in the School of Urban and Regional Planning at Ryerson University. Her work focuses on environmental planning, especially resilient design and ecology, and she has founded both a research center and a design practice that explore systems resilience.


Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite.

Nocturnal creatures generally have highly developed senses of hearing, smell, and specially adapted eyesight. Such traits can help animals such as the Helicoverpa zea moths avoid predators. Some animals, such as cats and ferrets, have eyes that can adapt to both low-level and bright day levels of illumination (see metaturnal). Others, such as bushbabies and (some) bats, can function only at night. Many nocturnal creatures including tarsiers and some owls have large eyes in comparison with their body size to compensate for the lower light levels at night. More specifically, they have been found to have a larger cornea relative to their eye size than diurnal creatures to increase their visual sensitivity: in the low-light conditions. Nocturnality helps wasps, such as Apoica flavissima, avoid hunting in intense sunlight.

Diurnal animals, including squirrels and songbirds, are active during the daytime. Crepuscular species, such as rabbits, skunks, tigers, and hyenas, are often erroneously referred to as nocturnal. Cathemeral species, such as fossas and lions, are active both in the day and at night.

While most humans are diurnal, for various personal and social/cultural reasons some people are temporarily or habitually nocturnal.

The most known creatures to be nocturnal include cats, rodents, and owls, which all have heightened senses (including their sense of sight).

North Cape oil spill

The North Cape oil spill took place on January 19, 1996, when the tank barge North Cape and the tug Scandia grounded on Moonstone Beach in South Kingstown, Rhode Island, after the tug caught fire in its engine room during a winter storm. An estimated 828,000 gallons of home heating oil was spilled. Oil spread throughout a large area of Block Island Sound, including Trustom Pond National Wildlife Refuge, resulting in the closure of a 250-square-mile (650 km2) area of the sound for fishing.

Hundreds of oiled birds and large numbers of dead lobsters, surf clams, and sea stars were recovered in the weeks following the spill. US federal and Rhode Island state governments undertook considerable work to clean up the spill and restore lost fishery stocks and coastal marine habitat. The North Cape oil spill is considered a significant legal precedent in that it was the first major oil spill in the continental U.S. after the passage of the Oil Pollution Act of 1990, resulting from the Exxon Valdez oil spill in Alaska on March 24, 1989.

Potential natural vegetation

In ecology, potential natural vegetation (PNV) is the vegetation that would be expected given environmental constraints (climate, geomorphology, geology) without human intervention or a hazard event.

The concept has been developed in the mid 1950s by phytosociologist Reinhold Tüxen, partly expanding on the concept of climax.

Resistance (ecology)

In the context of ecological stability, resistance is the property of communities or populations to remain "essentially unchanged" when subject to disturbance. The inverse of resistance is sensitivity.

Saeid Eslamian

Saeid Eslamian is a Full Professor of Hydrology and Water Resources Sustainability at Isfahan University of Technology in the Department of Water Engineering. His research focuses mainly on Statistical and Environmental Hydrology and Climate Change. In particular, he is working on Modeling Natural Hazards including Flood, Drought, Storm, Wind, Pollution toward a sustainable environment.

Formerly, he was a Visiting Professor at Princeton University, United States, University of ETH Zurich, Switzerland and McGill University, Montreal, Quebec, Canada.

He has contributed to more than 400 publications in journals, books, or as technical reports. He is the Founder and Chief Editor of both International Journal of Hydrology Science and Technology and Journal of Flood Engineering.

Currently, he has been the author of about 100 Books and Chapters.

Prof. Eslamian is the editorial board member and reviewer of about 40 Web of Science (ISI) Journals including ASCE Journal of Hydrologic Engineering, ASCE Journal of Water Resources Planning and Management.

He is a member of the following associations: American Society of Civil Engineers (ASCE), International Association of Hydrologic Science (IAHS), World Conservation Union (IUCN), GC Network for Drylands Research and Development (NDRD), International Association for Urban Climate (IAUC), International Society for Agricultural Meteorology, UK Chinese Association of Resources and Environment, Association of Water and Environment Modeling.From 2010, Professor has started scientific collaboration internationally toward Sustainable Development and Ecological resilience. He is currently the head of International Collaboration for Environmental Sustainability (ICES) having about 1000 members including Professors, Directors, Sustainability Professionals, Policy Makers, Senior and Graduate Students.

Sustainability science

Sustainability science emerged in the 21st century as a new academic discipline. This new field of science was officially introduced with a "Birth Statement" at the World Congress "Challenges of a Changing Earth 2001" in Amsterdam organized by the International Council for Science (ICSU), the International Geosphere-Biosphere Programme (IGBP), the International Human Dimensions Programme on Global Environmental Change and the World Climate Research Programme (WCRP).

The field reflects a desire to give the generalities and broad-based approach of “sustainability” a stronger analytic and scientific underpinning as it "brings together scholarship and practice, global and local perspectives from north and south, and disciplines across the natural and social sciences, engineering, and medicine". Ecologist William C. Clark proposes that it can be usefully thought of as "neither 'basic' nor 'applied' research but as a field defined by the problems it addresses rather than by the disciplines it employs" and that it "serves the need for advancing both knowledge and action by creating a dynamic bridge between the two".The field is focused on examining the interactions between human, environmental, and engineered systems to understand and contribute to solutions for complex challenges that threaten the future of humanity and the integrity of the life support systems of the planet, such as climate change, biodiversity loss, pollution and land and water degradation.Sustainability science, like sustainability itself, derives some impetus from the concepts of sustainable development and environmental science. Sustainability science provides a critical framework for sustainability while sustainability measurement provides the evidence-based quantitative data needed to guide sustainability governance.

Teton River (Idaho)

The Teton River is an 64-mile-long (103 km) tributary of the Henrys Fork of the Snake River in southeastern Idaho in the United States. It drains through the Teton Valley along the west side of the Teton Range along the Idaho-Wyoming border at the eastern end of the Snake River Plain. Its location along the western flank of the Tetons provides the river with more rainfall than many other rivers of the region.

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