Chris Perrins

Christopher Miles Perrins, LVO FRS[5] (born 11 May 1935)[1][6] is Emeritus Fellow of the Edward Grey Institute of Field Ornithology at the University of Oxford, Emeritus Fellow at Wolfson College, Oxford[7] and Her Majesty's Warden of the Swans since 1993.[1][7][8]

Chris Perrins

Born11 May 1935 (age 84)[1]
EducationCharterhouse School
Alma mater
Mary Ceresole Carslake (m. 1963)
AwardsGodman-Salvin Medal
Scientific career
ThesisSome factors influencing brood-size and populations in tits (1963)
Doctoral advisorDavid Lack[2]
Doctoral students


Perrins was educated at Charterhouse School and Queen Mary College[1] where he was awarded a Bachelor of Science degree in Zoology in 1957.[1] He completed his postgraduate study and research at the University of Oxford where he was awarded a Doctor of Philosophy degree in 1963 for research on brood size in tits supervised by David Lack.[2]

Research and career

Perrins research interests are in the population dynamics and breeding biology of birds, particularly tits (Paridae),[2][9] mute swans[10][11] and seabirds on Skomer and Skokholm.[7][12][13] He investigated animal lead poisoning of swans from lead shot.[12] He is renowned for his work on avian population ecology and, in particular, reproductive rates. He has made a number of important contributions to the long-term study of the great tit at Wytham Woods[14] — an area of mixed woodland established in 1947 by evolutionary biologist David Lack – one of the most famous studies in population ecology.[5]

He was the first to discover that avian clutch size – the number of eggs laid in a single nesting – in great tits has a remarkably high heritability and that the likelihood of the survival of young birds can be traced back to nutrition in the nest.[5] Perrins also demonstrated that females lay a clutch of an appropriate size for their ability to feed.[5][15] He supervised several successful DPhil students at Oxford including Matt Ridley[4] and Tim Birkhead.[3]

According to Scopus,[16] As of 2016 his most cited journal articles have been published in Ibis,[17][18] Nature,[19] Science[9] and the Journal of Animal Ecology.[20]


  • British Tits[21]
  • Encyclopedia of Birds[22]
  • The Mute Swan[23]
  • The Illustrated Encyclopedia of Birds: The Definitive Guide to Birds of the World: Christopher M. Perrins Hardline (1990) ISBN 978-0747202776
  • Bird Population Studies: Relevance to Conservation and Management: (Oxford Ornithology Series) by Christopher M. Perrins, G. J. Hirons and J. D. Lebreton (1996) ISBN 978-0198540823
  • The Birds of the Western Palearctic, Volume 1: Non-passerines David Snow, Christopher M. Perrins and Robert Gillmor (1998) Oxford University Press. ISBN 978-0198501879
  • The Birds of the Western Palearctic, Volume 2: by David Snow, Christopher M. Perrins and Robert Gillmor (1998) Oxford University Press. ISBN 978-0198501886
  • The New Encyclopedia of Birds: Editor, (2003) Oxford University Press. ISBN 978-0198525066
  • The Complete Encyclopedia of Birds and Bird Migration: Jonathan Elphick: Christopher M. Perrins (2004) ISBN 978-0785816676

Awards and honours

Perrins has received a number of awards for his research, including the Godman-Salvin Medal of the British Ornithologists' Union in 1988, and the Royal Society for the Protection of Birds (RSPB) Medal in 1992. In 1993, he was appointed as the first Warden of the Swans in the Royal Household, playing an important role in the annual Swan Upping ceremony. This was a new office in the Royal Household of the Sovereign of the United Kingdom, created in 1993. Other awards and honours include:[7]


  1. ^ a b c d e f g h Anon (1992). Perrins, Prof. Christopher Miles. Who's Who (online Oxford University Press ed.). A & C Black, an imprint of Bloomsbury Publishing plc. doi:10.1093/ww/9780199540884.013.30593. (subscription or UK public library membership required) (subscription required)
  2. ^ a b c Perrins, Christopher Miles (1963). Some factors influencing brood-size and populations in tits (DPhil thesis). University of Oxford. OCLC 44835614. EThOS
  3. ^ a b Birkhead, Timothy Robert (1976). Breeding biology and survival of guillemots (Uria aalge). (DPhil thesis). University of Oxford. OCLC 44837387. EThOS Free to read.
  4. ^ a b Ridley, Matthew White (1983). Mating system of the pheasant (Phasianus colchicus) (DPhil thesis). University of Oxford. OCLC 52225811. EThOS
  5. ^ a b c d e Anon (1997). "Professor Christopher Perrins LVO FRS". London: Archived from the original on 17 November 2015. One or more of the preceding sentences incorporates text from the website where:

    "All text published under the heading 'Biography' on Fellow profile pages is available under Creative Commons Attribution 4.0 International License." --"Royal Society Terms, conditions and policies". Archived from the original on 25 September 2015. Retrieved 9 March 2016.CS1 maint: BOT: original-url status unknown (link)

  6. ^ "Weekend birthdays", The Guardian, p. 52, 10 May 2014
  7. ^ a b c d Perrins, Chris (2015). "Professor Chris Perrins LVO FRS". Oxford: University of Oxford. Archived from the original on 28 March 2015.
  8. ^ Christopher Perrins, Professor of Zoology at Oxford University talks about Swan Upping on YouTube
  9. ^ a b Charmantier, A.; McCleery, R. H.; Cole, L. R.; Perrins, C.; Kruuk, L. E. B.; Sheldon, B. C. (2008). "Adaptive Phenotypic Plasticity in Response to Climate Change in a Wild Bird Population". Science. 320 (5877): 800–803. doi:10.1126/science.1157174. PMID 18467590.
  10. ^ Charmantier, A.; Perrins, C.; McCleery, R. H.; Sheldon, B. C. (2006). "Quantitative genetics of age at reproduction in wild swans: Support for antagonistic pleiotropy models of senescence". Proceedings of the National Academy of Sciences. 103 (17): 6587–6592. doi:10.1073/pnas.0511123103. PMC 1458927. PMID 16618935. open access
  11. ^ Mccleery, R. H.; Perrins, C. M.; Wheeler, D.; Groves, S. (2006). "The effect of breeding status on the timing of moult in Mute Swans Cygnus olor". Ibis. 149 (1): 86–90. doi:10.1111/j.1474-919X.2006.00596.x.
  12. ^ a b Perrins, Chris (2016). "Oxford Alumni: Professor Chris Perrins". Oxford: University of Oxford. Archived from the original on 21 July 2016.
  13. ^ Ropert-Coudert, Yan; Guilford, Tim; Freeman, Robin; Boyle, Dave; Dean, Ben; Kirk, Holly; Phillips, Richard; Perrins, Chris (2011). "A Dispersive Migration in the Atlantic Puffin and Its Implications for Migratory Navigation". PLOS ONE. 6 (7): e21336. doi:10.1371/journal.pone.0021336. PMC 3140476. PMID 21799734. open access
  14. ^ Savill, Peter; Perrins, Christopher; Kirby, Keith; Fisher, Nigel (2011). Wytham Woods: Oxford's Ecological Laboratory. Oxford: Oxford University Press. p. 282. doi:10.1093/acprof:osobl/9780199605187.001.0001. ISBN 9780199605187. Archived from the original on 23 August 2016.
  15. ^ Dean, B.; Freeman, R.; Kirk, H.; Leonard, K.; Phillips, R. A.; Perrins, C. M.; Guilford, T. (2012). "Behavioural mapping of a pelagic seabird: combining multiple sensors and a hidden Markov model reveals the distribution of at-sea behaviour". Journal of the Royal Society Interface. 10 (78): 20120570–20120570. doi:10.1098/rsif.2012.0570. PMC 3565783. PMID 23034356. open access
  16. ^ Chris Perrins's publications indexed by the Scopus bibliographic database. (subscription required)
  17. ^ Perrins, C. M. (2008). "Tits and their caterpillar food supply". Ibis. 133: 49–54. doi:10.1111/j.1474-919X.1991.tb07668.x.
  18. ^ Perrins, C. M. (2008). "Eggs, egg formation and the timing of breeding". Ibis. 138 (1): 2–15. doi:10.1111/j.1474-919X.1996.tb04308.x.
  19. ^ Pettifor, R. A.; Perrins, C. M.; McCleery, R. H. (1988). "Individual optimization of clutch size in great tits". Nature. 336 (6195): 160–162. doi:10.1038/336160a0.
  20. ^ Noordwijk, A.J. Van; McCleery, R.H.; Perrins, C.M. (1995). "Selection for the Timing of Great Tit Breeding in Relation to Caterpillar Growth and Temperature". The Journal of Animal Ecology. 64 (4): 451. doi:10.2307/5648.
  21. ^ British Tits (1979) Collins New Naturalist Series ISBN 978-0002195379
  22. ^ Encyclopedia of Birds (1985) Christopher M. Perrins and A.L.A. Middleton, Facts on File Inc ISBN 978-0816011506
  23. ^ The Mute Swan (1986) Helm Field Guides, Birkhead M. Perrins C. ISBN 978-0709932598
1935 in the United Kingdom

Events from the year 1935 in the United Kingdom. This year was the Silver Jubilee of King George V. Political events include a general election in November and changes in the leadership of both the Conservative and Labour parties.

2015 CrossFit Games

The 2015 CrossFit Games were held on July 21–26, 2015 at the StubHub Center in Carson, California. Ben Smith, of Virginia, was the men's winner. A Games competitor every year since 2009, this was his first win (he placed third in 2011 and 2013). Smith secured the win in a dramatic final event by holding off 2014's second-place finisher Mat Fraser who many had predicted would be the 2015 champion. Iceland's Katrin Davidsdottir, making her third Games appearance, was the women's winner, overtaking countrywoman Ragnheiður Sara Sigmundsdottir, who led for much of the competition, in the final event. CrossFit Mayhem Freedom from Cookeville, Tennessee, captained by four-time individual men's champion Rich Froning Jr., won the Affiliate Cup.

Approximately 273,000 participants signed up for the 2015 CrossFit Games Open, which was held over the course of five weeks in February and March. The top 20 men, 20 women, and 15 teams from the 12 American and Canadian regions, the top 30 men, 30 women, and 20 teams in the European and Australian regions, and the top 10 men, 10 women, and 10 teams from Latin America, Africa, and Asia qualified for the CrossFit Regionals. After three-day Regional competitions held around the world, the field advancing to the CrossFit Games was narrowed to 40 men, 40 women, and 40 teams. New for the 2015 Regionals, the 17 regions were combined into 8 "super-regions".As in previous years, the men's and women's competitions were similar, with the only differences being weights of lifted objects, heights of obstacles, etc. First-time elements in 2015 included open-ocean paddleboard racing and pegboard climbing.

CrossFit also held a Masters competition where athletes aged 40 and above competed in their own age groups. Masters athletes qualified though the Open followed by an online Games qualification, called the Masters Qualifier, instead of Regionals. The top 20 Masters men and women in each age group were invited to participate at the Games.

In 2015, the CrossFit Games added a Teenage competition to the Open in two age groups (14-15 and 16-17) for boys and girls. Towards the end of the Open, it was announced that the top 10 in each age group after the completion of the Open would qualify to participate in the Games. The Teenage and Masters Games events were the same and took place from July 21–23.

Alfred Newton Lecture

The Alfred Newton Lecture is an academic prize lecture awarded by the British Ornithological Union. It is named for Alfred Newton.

David Lack

David Lambert Lack FRS (16 July 1910 – 12 March 1973) was a British evolutionary biologist who made contributions to ornithology, ecology, and ethology. His 1947 book, Darwin's Finches, on the finches of the Galapagos Islands was a landmark work as were his other popular science books on Life of the Robin and Swifts in a Tower. He developed what is now known as Lack's Principle which explained the evolution of avian clutch sizes in terms of individual selection as opposed to the competing contemporary idea that they had evolved for the benefit of species (also known as group selection). His pioneering life-history studies of the living bird helped in changing the nature of ornithology from what was then a collection-oriented field. He was a longtime director of the Edward Grey Institute of Field Ornithology at the University of Oxford.

Department of Zoology, University of Oxford

The Department of Zoology is the zoology department of the University of Oxford, England, which is part of the university's Mathematical, Physical and Life Sciences Division.

Edward Grey Institute of Field Ornithology

The Edward Grey Institute of Field Ornithology (EGI), at Oxford University in England, is an academic body that conducts research in ornithology and the general field of evolutionary ecology and conservation biology, with an emphasis on understanding organisms in natural environments. It is named in honour of Edward Grey, 1st Viscount Grey of Fallodon, a notable politician and ornithologist, and is part of the Department of Zoology at Oxford University.

The Institute houses the Alexander Library, the best ornithological library in Europe and one of the best in the world, which is named after W. B. Alexander.

List of Old Carthusians

The following are notable Old Carthusians, who are former pupils of Charterhouse (founded in 1611).

Matt Ridley

Matthew White Ridley, 5th Viscount Ridley (born 7 February 1958), commonly known as Matt Ridley, is a British journalist and businessman. Ridley is best known for his writings on science, the environment, and economics. He has written several science books including The Red Queen: Sex and the Evolution of Human Nature (1994), Genome (1999), The Rational Optimist: How Prosperity Evolves (2010) and The Evolution of Everything: How Ideas Emerge (2015). He publishes a blog, and has been a regular contributor to The Times newspaper.

Ridley is a libertarian, and a staunch supporter of Brexit. Since 2013, he has been a Conservative hereditary peer, with a seat in the House of Lords.Ridley was chairman of the UK bank Northern Rock from 2004 to 2007, during which period Northern Rock experienced the first run on a British bank in 150 years. Ridley resigned and the bank was bailed out by the UK government leading to the nationalisation of Northern Rock.

New Naturalist

The New Naturalist Library (also known as The New Naturalists) is a series of books published by Collins in the United Kingdom, on a variety of natural history topics relevant to the British Isles. The aim of the series at the start was: "To interest the general reader in the wild life of Britain by recapturing the inquiring spirit of the old naturalists." An editors' preface to a 1952 monograph says: "An object of the New Naturalist series is the recognition of the many-sidedness of British natural history, and the encouragement of unusual and original developments of its forgotten or neglected facets."The first volume to appear was E.B. Ford's Butterflies in 1945. The authors of this series are usually eminent experts, often professional scientists. This gives the series authority, and many are or have been authoritative introductory textbooks on a subject for some years. The books are written in scientific style, but are intended to be readable by the non-specialist, and are an early example of popular science in the media.

The books of the series have had considerable influence on many students who later became professional biologists, such as W.D. Hamilton and Mike Majerus. The latter was inspired by Ford's Butterflies and Moths, and has since added two volumes of his own to the series.

A parallel series known as the New Naturalist Monograph Library (and often referred to as The New Naturalist Special Volumes) was also published. Its aim was to cover "in greater detail... a single species or group of species". There have been no additions to the Monograph Library since 1971.

Volume 82 of the main series, The New Naturalists, described the series to date, with authors' biographies and a guide to collecting the books.

The original Editorial Board consisted of Julian Huxley, James Fisher, Dudley Stamp, John Gilmour and Eric Hosking. Until 1985, the highly characteristic dust jacket illustrations were by Rosemary and Clifford Ellis; since then they have been by Robert Gillmor.

Being a numbered series, with a very low print run for some volumes, the books are highly collectable. Second-hand copies of the rarer volumes, in good condition, can command high prices. The 100th volume, Woodlands by Oliver Rackham was published in 2006. Woodlands (volume 100) was also published in 2006 as a "leatherbound" edition, limited to 100 copies. In fact it was fake leather. The second "leatherbound" New Naturalist - Dragonflies by Philip Corbet and Stephen Brooks - was published in 2008. The (fake) leather edition of Dragonflies (volume 106) was initially limited to 400 copies, which was subsequently limited to 303, and finally to 250. According to the New Naturalist website only 217 were actually sold and the remaining unsold stock is being kept secure at HarperCollins's offices. HarperCollins continue to produce limited numbers of "leatherbound" editions of all volumes published since Dragonflies, but only from Islands (volume 109) was real leather actually used. All recent volumes have only 50 leatherbound copies.

The series won the 2007 British Book Design and Production Award for "brand or series identity", and in 2008 the official website was launched, with features including the latest news, a members only area with access to exclusive content and downloads, and a forum.

In around 1990, Bloomsbury produced a series of facsimile editions, as hardbacks with new dustjacket designs, and with all plates in black and white, including those which were originally in colour.

Pat Nuttall

Patricia (Pat) Anne Nuttall, OBE (born 1953) is a British virologist and acarologist known for her research on tick-borne diseases. Her discoveries include the fact that pathogens can be transmitted between vectors feeding on a host without being detectable in the host's blood. She is also a science administrator who served as the director of the Natural Environment Research Council (NERC) Centre for Ecology & Hydrology (2001–11). As of 2015, she is professor of arbovirology in the Department of Zoology of the University of Oxford.


Perrins may refer to:

Charles William Dyson Perrins (1864–1958), English businessman, bibliophile and philanthropist

Chris Perrins, LVO, FRS (born 1935), British biologist

George Perrins (1873 – unknown) was an English footballer

Isaac Perrins, English bareknuckle prizefighter and 18th-century engineer

Leslie Perrins (1901–1962), English actor who often played villains

Wesley Perrins, MBE (1905–1990), English trade unionist and Labour Party politician from Stourbridge

William Henry Perrins (1793–1867), drug-store chemist who formed a partnership in 1823 with John Wheeley Lea

RSPB Medal

The RSPB Medal is awarded annually by the Royal Society for the Protection of BirdsAccording to the RSPB:

The RSPB Medal is the Society's most prestigious award. It is presented to an individual in recognition of wild bird protection and countryside conservation. It is usually awarded annually to one or occasionally two people.

The medal was first awarded in 1908.

Tim Birkhead

Timothy Robert Birkhead (born 1950) is a British ornithologist. He has been Professor of Behaviour and Evolution at the University of Sheffield since 1976.

Witherby Memorial Lecture

The Witherby Memorial Lecture is an academic lectureship awarded by the British Trust for Ornithology (BTO) annually since 1968.


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