Centipedes (from Latin prefix centi-, "hundred", and pes, pedis, "foot") are predatory arthropods belonging to the class Chilopoda of the subphylum Myriapoda, an arthropod group which also includes Millipedes and other multi-legged creatures. Centipedes are elongated metameric creatures with one pair of legs per body segment. Most centipedes are generally venomous and could inflict a painful bite, injecting their venom through pincer-like appendage known as forcipules. Despite the name, centipedes can have a varying number of legs, ranging from 30 to 354. Centipedes always have an odd number of pairs of legs. Therefore, no centipede has exactly 100 legs. Similar to spiders and scorpions, centipedes are predominantly carnivorous.:168
Their size can range from a few millimetres in the smaller lithobiomorphs and geophilomorphs to about 30 cm (12 in) in the largest scolopendromorphs. Centipedes can be found in a wide variety of environments. They normally have a drab coloration combining shades of brown and red. Cavernicolous (cave-dwelling) and subterranean species may lack pigmentation, and many tropical scolopendromorphs have bright aposematic colors.
Worldwide, an estimated 8,000 species of centipedes are thought to exist, of which 3,000 have been described. Centipedes have a wide geographical range, where they even reach beyond the Arctic Circle. They are found in an array of terrestrial habitats from tropical rainforests to deserts. Within these habitats, centipedes require a moist microhabitat because they lack the waxy cuticle of insects and arachnids, therefore causing them to rapidly lose water. Accordingly, they are found in soil and leaf litter, under stones and dead wood, and inside logs. Centipedes are among the largest terrestrial invertebrate predators, and often contribute significantly to the invertebrate predatory biomass in terrestrial ecosystems.
|Orders and Families|
Centipedes have a rounded or flattened head, bearing a pair of antennae at the forward margin. They have a pair of elongated mandibles, and two pairs of maxillae. The first pair of maxillae form the lower lip, and bear short palps. The first pair of limbs stretch forward from the body to cover the remainder of the mouth. These limbs, or maxillipeds, end in sharp claws and include venom glands that help the animal to kill or paralyze its prey.
Many species of centipedes lack eyes, but some possess a variable number of ocelli, which are sometimes clustered together to form true compound eyes. However, these eyes are only capable of discerning light and dark, and have no true vision. In some species, the first pair of legs at the head end of the centipede acts as sense organs similar to antennae, but unlike the antennae of most other animals, theirs point backwards. Unusual sense organs found in some groups are the organs of Tömösváry. These are located at the base of the antennae, and consist of a disc-like structure with a central pore surrounded by sensory cells. They are probably used for sensing vibrations, and may even provide a sense of hearing.
Forcipules are a unique feature found only in centipedes and in no other arthropods. The forcipules are modifications of the first pair of legs, forming a pincer-like appendage always found just behind the head. Forcipules are not true mouthparts, although they are used in the capture of prey items, injecting venom and holding onto captured prey. Venom glands run through a tube almost to the tip of each forcipule.
Behind the head, the body consists of 15 or more segments. Most of the segments bear a single pair of legs, with the maxillipeds projecting forward from the first body segment, and the final two segments being small and legless. Each pair of legs is slightly longer than the pair immediately in front of it, ensuring that they do not overlap, so reducing the chance that they will collide with each other while moving swiftly. In extreme cases, the last pair of legs may be twice the length of the first pair. The final segment bears a telson and includes the openings of the reproductive organs.
As predators, centipedes mainly use their antennae to seek out their prey. The digestive tract forms a simple tube, with digestive glands attached to the mouthparts. Like insects, centipedes breathe through a tracheal system, typically with a single opening, or spiracle on each body segment. They excrete waste through a single pair of malpighian tubules.
Scolopendra gigantea, also known as the Amazonian giant centipede, is the largest existing species of centipede in the world, reaching over 30 cm (12 in) in length. It is known to eat lizards, frogs, birds, mice, and even bats, catching them in midflight, as well as rodents and spiders.
Centipede reproduction does not involve copulation. Males deposit a spermatophore for the female to take up. In one clade, this spermatophore is deposited in a web, and the male undertakes a courtship dance to encourage the female to engulf his sperm. In other cases, the males just leave them for the females to find. In temperate areas, egg laying occurs in spring and summer, but in subtropical and tropical areas, little seasonality to centipede breeding is apparent. A few species of parthenogenetic centipedes are known.
The Lithobiomorpha and Scutigeromorpha lay their eggs singly in holes in the soil, and the female fills the holes with soil and leaves them. The number of eggs laid ranges from about 10 to 50. Time of development of the embryo to hatching is highly variable and may take from one to a few months. Time of development to reproductive period is highly variable within and among species. For example, it can take 3 years for S. coleoptrata to achieve adulthood, whereas under the right conditions, lithobiomorph species may reach a reproductive period in 1 year. In addition, centipedes are relatively long-lived when compared to insects. For example, the European Lithobius forficatus may live for 5 to 6 years, and the wide-ranging Scolopendra subspinipes can live for over 10 years. The combination of a small number of eggs laid, long gestation period, and long time of development to reproduction has led authors to label lithobiomorph centipedes as K-selected.
Females of the Geophilomorpha and Scolopendromorpha show far more parental care. The eggs, 15 to 60 in number, are laid in a nest in the soil or in rotten wood. The female stays with the eggs, guarding and licking them to protect them from fungi. The female in some species stays with the young after they have hatched, guarding them until they are ready to leave. If disturbed, the female either abandons the eggs or eats them; abandoned eggs tend to fall prey to fungi rapidly. Some species of Scolopendromorpha are matriphagic, meaning the offspring eat their mother.
Little is known of the life history of the Craterostigmomorpha.
Centipedes grow their legs at different points in their development. In the primitive condition, exhibited by the Lithobiomorpha, Scutigeromorpha, and Craterostigmomorpha, development is anamorphic: more pairs of legs are grown between moults. For example, Scutigera coleoptrata, the American house centipede, hatches with only four pairs of legs and in successive moults has 5, 7, 9, 11, 15, 15, 15 and 15 before becoming a sexually mature adult. Life stages with fewer than 15 pairs of legs are called larval stadia (about five stages). After the full complement of legs is achieved, the now postlarval stadia (about five stages) develop gonopods, sensory pores, more antennal segments, and more ocelli. All mature lithobiomorph centipedes have 15 leg-bearing segments.:27
The Craterostigmomorpha only have one phase of anamorphosis, with embryos having 12 pairs, and moultees 15.
The clade Epimorpha, consisting of the orders Geophilomorpha and Scolopendromorpha, exhibits epimorphy: all pairs of legs are developed in the embryonic stages, and offspring do not develop more legs between moults. This clade contains the longest centipedes; the maximum number of thoracic segments may also vary intraspecifically, often on a geographical basis; in most cases, females bear more legs than males. The number of leg-bearing segments varies widely, from 15 to 191, but the developmental mode of their creation means they are always added in pairs—hence the total number of pairs is always odd.
Centipedes are predominantly generalist predators, which means they have adapted to eat a variety of different available prey. Examination of centipede gut contents suggests that plant material is an unimportant part of their diets, although centipedes have been observed to eat vegetable matter when starved during laboratory experiments.:168
Centipedes are mostly nocturnal. Studies on their activity rhythms confirm this, although a few observations of centipedes active during the day have been made, and one species, Strigamia chinophila, is diurnal. What centipedes actually eat is not well known because of their cryptic lifestyles and thorough mastication of food. Laboratory feeding trials support that they will feed as generalists, taking almost anything that is soft-bodied and in a reasonable size range. Earthworms may provide the bulk of diets for geophilomorphs, since they burrow through the soil and earthworm bodies would be easily pierced by their venom claws. Geophilomorphs probably cannot subdue earthworms larger than themselves, so smaller earthworms may be a substantial proportion of their diet.
Scolopendromorphs, given their size, are able to feed on vertebrates, in addition to invertebrates. They have been observed eating reptiles, amphibians, small mammals, bats, and birds. Springtails may provide a large proportion of lithobiomorph diets. Little is known about scutigeromorph or craterostigmomorph diets. All centipedes are potential intraguild predators. Centipedes and spiders may frequently prey on one another. Two species, Scolopendra cataracta and S. paradoxa are known to be amphibious and are believed to hunt aquatic or amphibious invertebrates.
Many larger animals prey upon centipedes, such as mongooses, mice, salamanders, beetles and snakes.:354–356 They form an important item of diet for many species and the staple diet of some such as the African ant Amblyopone pluto, which feeds solely on geophilomorph centipedes, and the South African Cape black-headed snake Aparallactus capensis.:354–356
Centipede defences include their speed and venomous forcipules, as well as the secretion of defensive chemicals. Geophilomorph centipedes can secrete sticky substances that generate toxic hydrogen cyanide and benzoic acid from microscopic glands on their undersides. Similarly, lithobiomorph centipedes secrete a sticky substance from glands in the rear-most two pairs of legs.
Water regulation is an important aspect of centipede ecology, since they lose water rapidly in dry conditions and are found in moist microhabitats. Water loss is a result of centipedes lacking a waxy covering of their exoskeleton and excreting waste nitrogen as ammonia, which requires extra water. Centipedes deal with water loss through a variety of adaptations. Geophilomorphs lose water less rapidly than lithobiomorphs, though they have a greater surface area to volume ratio. This may be because geophilomorphs have a more heavily sclerotized pleural membrane. Spiracle shape, size, and ability to constrict also have an influence on rate of water loss. In addition, the number and size of coxal pores may be variables affecting centipede water balance.
Centipedes live in many different habitat types—forest, savannah, prairie, and desert, to name a few. Some geophilomorphs are adapted to littoral habitats, where they feed on barnacles. Species of all orders excluding the Craterostigmomorpha have adapted to caves. Centipede densities have been recorded as high as 600/m2 and biomass as high as 500 mg/m2 wet weight. Small geophilomorphs attain highest densities, followed by small lithobiomorphs. Large lithobiomorphs attain densities of 20/m2. One study of scolopendromorphs records Scolopendra morsitans in a Nigerian savannah at a density of 0.16/m2 and a biomass of 140 mg/m2 wet weight.
As a food item, certain large-sized centipedes are consumed in China, usually skewered and grilled or deep fried. They are often seen in street vendor's stalls in large cities, including Donghuamen and Wangfujing markets in Beijing.
Also in China, as well as in Laos, Thailand, and Cambodia, large centipedes are kept in liquor for a period of time. This custom is allegedly part of the traditional Chinese medicine. Said to have medicinal properties and to be reinvigorating, the liquor with the centipede submerged in it is consumed as a special drink.
Some species of centipedes can be hazardous to humans because of their bite. Although a bite to an adult human is usually very painful and may cause severe swelling, chills, fever, and weakness, it is unlikely to be fatal. Bites can be dangerous to small children and those with allergies to bee stings. The venomous bite of larger centipedes can induce anaphylactic shock in such people. Smaller centipedes are generally incapable of piercing human skin.
Even nonvenomous centipedes are considered frightening by humans due to their dozens of legs moving at the same time and their tendency to dart swiftly out of the darkness towards one's feet. A 19th-century Tibetan poet warned his fellow Buddhists, "if you enjoy frightening others, you will be reborn as a centipede."
|Internal phylogeny of the Chilopoda|
|The upper three groups form the paraphyletic Anamorpha.|
The fossil record of centipedes extends back to , during the Late Silurian. They belong to the subphylum Myriapoda which includes Diplopoda, Symphyla, and Pauropoda. The oldest known fossil land animal, Pneumodesmus newmani, is a myriapod. Being among the earliest terrestrial animals, centipedes were one of the first to fill a fundamental niche as ground level generalist predators in detrital food webs. Today, centipedes are abundant and exist in many harsh habitats.
Within the myriapods, centipedes are believed to be the first of the extant classes to branch from the last common ancestor. The five orders of centipedes are: Craterostigmomorpha, Geophilomorpha, Lithobiomorpha, Scolopendromorpha, and Scutigeromorpha. These orders are united into the clade Chilopoda by the following synapomorphies:
The Chilopoda are then split into two clades: the Notostigmophora including the Scutigeromorpha and the Pleurostigmophora including the other four orders. The main difference is that the Notostigmomorpha have their spiracles located mid-dorsally. It was previously believed that Chilopoda was split into Anamorpha (Lithobiomorpha and Scutigeromorpha) and Epimorpha (Geophilomorpha and Scolopendromorpha), based on developmental modes, with the relationship of the Craterostigmomorpha being uncertain. Recent phylogenetic analyses using combined molecular and morphological characters supports the previous phylogeny. The Epimorpha still exist as a monophyletic group within the Pleurostigmophora, but the Anamorpha are paraphyletic.
Geophilomorph centipedes have been used to argue for the developmental constraint of evolution; that the evolvability of a trait, the number of segments in the case of geophilomorph centipedes, was constrained by the mode of development. The geophilomorph centipedes have variable segment numbers within species, yet as with all centipedes, they always have an odd number of pairs of legs. In this taxon, the number of segments ranges from 27 to 191, but is never an even number.
The Scutigeromorpha are anamorphic, reaching 15 leg-bearing segments in length. Also known as house centipedes, they are very fast creatures, and able to withstand falling at great speed: they reach up to 15 body lengths per second when dropped, surviving the fall. They are the only centipede group to retain their original compound eyes, within which a crystalline layer analogous to that seen in chelicerates and insects can be observed. They also bear long and multi-segmented antennae. Adaptation to a burrowing lifestyle has led to the degeneration of compound eyes in other orders; this feature is of great use in phylogenetic analysis.
The group is the sole extant representative of the Notostigmomorpha, defined by having a single spiracle opening at the posterior of each dorsal plate. The more derived groups bear a plurality of spiracular openings on their sides, and are termed the Pleurostigmomorpha. Some even have several unpaired spiracles that can be found along the mid-dorsal line and closer to their posterior section of tergites. There are three families: Pselliodidae, Scutigeridae and Scutigerinidae. Pselliodidae includes just a few species in the genus Sphendononema (=Pselliodes), occurring in the Neotropics and tropical Africa. Scutigerinidae, composed of three species in the genus Scutigerina, is restricted to southern Africa and Madagascar. Most scutigeromorphs from other parts of the world belong to the Scutigeridae, which includes two subfamilies, the Scutigerinae and Thereuoneminae.
The Lithobiomorpha, also known as stone centipedes, represent the other main group of anamorphic centipedes; they also reach a mature segment count of 15 trunk segments. This group has lost the compound eyes, and sometimes has no eyes altogether. Instead, its eyes have a single ocellus or a group of ocelli. Its spiracles are paired and can be found laterally. Every leg-bearing segment of this organism has a separate tergite, these alternating in length apart from a pair of long tergites on each of segments 7 and 8. It also has relatively short antennae and legs compared to the Scutigeromorpha. Two families are included, the Henicopidae and Lithobiidae.
The Craterostigmomorpha are the least diverse centipede clade, comprising only two extant species, both in the genus Craterostigmus. Their geographic range is restricted to Tasmania and New Zealand. They have a distinct body plan; their anamorphosis comprises a single stage: in their first moult, they grow from having 12 segments to having 15. Their low diversity and intermediate position between the primitive anamorphic centipedes and the derived Epimorpha has led to them being likened to the platypus. They represent the survivors of a once diverse clade.
Maternal brooding unites the Craterostigomomorpha with the Epimorpha into the clade Phylactometria. This trait is thought to be closely linked with the presence of sternal pores, which secrete sticky or noxious secretions, which mainly serve to repel predators and parasites. The presence of these pores on the Devonian Devonobius permits its inclusion in this clade, allowing its divergence to be dated to .
The Scolopendromorpha, also known as tropical centipedes, possess 21 or 23 body segments (apart from a single species, Scolopendropsis duplicata, which has 39 or 43 segments) with the same number of paired legs. Their antennae have 17 or more segments. The eyes have a fixed number of four ocelli on each side in the family Scolopendridae and one ocellus per side in the genus Mimops (family Mimopidae), but other families are blind. The order comprises the five families Cryptopidae, Scolopendridae, Mimopidae, Scolopocryptopidae, and Plutoniumidae. The only 2 known amphibious centipedes, Scolopendra cataracta and Scolopendra paradoxa belong to this order.
The Geophilomorpha, commonly known as soil centipedes, bear upwards of 27 leg-bearing segments. They are eyeless and blind, and bear spiracles on all leg-bearing segments—in contrast to other groups, which usually bear them only on their 3rd, 5th, 8th, 10th, 12th, and 14th segments—a "mid-body break", accompanied by a change in tagmatic shape, occurring roughly at the interchange from odd to even segments. This group, the most diverse at 1260 species, also contains the largest and leggiest specimens at 27 or more pairs of legs. They also have 14–segmented antennae. The group includes at least seven families: Mecistocephalidae, Geophilidae (including the former Linotaeniidae, Dignathodontidae and Macronicophilidae), Oryidae, Himantariidae, Schendylidae (including the former Ballophilidae), Zelanophilidae, and Gonibregmatidae (including the former Neogeophilidae and Eriphantidae).
|Scientific name||Common name|
|Alipes grandidieri||feather-tail centipede|
|Ethmostigmus trigonopodus||blue ring centipede|
|Lithobius forficatus||stone centipede|
|Pachymerium ferrugineum||earth centipede|
|Scolopendra galapagoensis||Galápagos centipede|
|Scolopendra cataracta||Aquatic centipede|
|Scutigera coleoptrata||house centipede|
|Scolopendra gigantea||Peruvian giant yellow-leg centipede|
|Scolopendra heros||giant red-headed centipede|
|Scolopendra morsitans||red-headed centipede|
|Scolopendra polymorpha||giant Sonoran centipede|
|Scolopendra subspinipes||Vietnamese centipede|
In 1902, C.L. Marlatt, an entomologist with the United States Department of Agriculture writes in Circular #48 - The House Centipede: It may often be seen darting across floors with great speed, occasionally stopping suddenly and remaining absolutely motionless, presently to resume its rapid movements, often darting directly at residents, particularly women, evidently with a desire to conceal itself beneath their dresses, creating much consternation.
"Centipede" is the debut single by American singer Rebbie Jackson and the title track from her debut album, Centipede.Centipede (band)
Centipede were an English jazz/progressive rock/big band with more than 50 members, organized and led by the British free jazz pianist Keith Tippett. Formed in 1970, it brought together much of a generation of young British jazz and rock musicians from a number of bands, including Soft Machine, King Crimson, Nucleus and Blossom Toes.Centipede performed several concerts in England, toured France, and recorded a double-album, Septober Energy (produced by Robert Fripp), before disbanding at the end of 1971. They reformed briefly in 1975 to play at a few French jazz festivals.Centipede (video game)
Centipede is a vertically oriented fixed shooter arcade game produced by Atari, Inc. in June 1981. The game was designed by Dona Bailey and Ed Logg. It was one of the most commercially successful games from the video arcade's golden age. The player fights off centipedes, spiders, scorpions and fleas, completing a round after eliminating the centipede that winds down the playing field.
Centipede was ported to Atari's own Atari 2600, Atari 5200, Atari 7800, and Atari 8-bit family. Under the Atarisoft label, the game was sold for the Apple II, Commodore 64, ColecoVision, VIC-20, IBM PC (as a booter), Intellivision, and TI-99/4A. Superior Software published the port for the BBC Micro.Centipede bite
A centipede bite is an injury resulting from the action of a centipede's forcipules, stinger-like appendages that pierce the skin and inject venom into the wound. Such a wound is not strictly speaking a bite, as the forcipules are a modified first pair of legs rather than true mouthparts. Clinically, the wound is viewed as a cutaneous condition characterized by paired hemorrhagic marks that form a chevron shape caused by the paired forcipules.The centipede's venom causes pain and swelling in the area of the bite, and may cause other reactions throughout the body. The majority of bites are not life-threatening to humans and present the greatest risk to children and those who develop allergic reactions.Centipede game
In game theory, the centipede game, first introduced by Robert Rosenthal in 1981, is an extensive form game in which two players take turns choosing either to take a slightly larger share of an increasing pot, or to pass the pot to the other player. The payoffs are arranged so that if one passes the pot to one's opponent and the opponent takes the pot on the next round, one receives slightly less than if one had taken the pot on this round. Although the traditional centipede game had a limit of 100 rounds (hence the name), any game with this structure but a different number of rounds is called a centipede game.
The unique subgame perfect equilibrium (and every Nash equilibrium) of these games indicates that the first player take the pot on the very first round of the game; however, in empirical tests, relatively few players do so, and as a result, achieve a higher payoff than the payoff predicted by the equilibria analysis. These results are taken to show that subgame perfect equilibria and Nash equilibria fail to predict human play in some circumstances. The Centipede game is commonly used in introductory game theory courses and texts to highlight the concept of backward induction and the iterated elimination of dominated strategies, which show a standard way of providing a solution to the game.HMAS Curlew
HMAS Curlew (M 1121) was a Ton-class minesweeper operated by the Royal Navy (as HMS Chediston) from 1953 to 1961, and the Royal Australian Navy from 1962 to 1991. During her Australian service, the ship operated off Malaysia during the Indonesia–Malaysia confrontation during the mid-1960s, then was modified for use as a minehunter. Delays in bringing a replacement class into service kept Curlew operational until 1990, and she was sold into civilian service in 1991.House centipede
A number of different centipede species in the family Scutigeridae are known as the house centipede, including:
Scutigera coleoptrata, originally from the Mediterranean region, but now found almost worldwide
Allothereua maculata, endemic to AustraliaJames and the Giant Peach
James and the Giant Peach is a popular children's novel written in 1961 by British author Roald Dahl. The original first edition published by Alfred Knopf featured illustrations by Nancy Ekholm Burkert. There have been reillustrated versions of it over the years, done by Michael Simeon for the first British edition, Emma Chichester Clark, Lane Smith and Quentin Blake. It was adapted into a film of the same name in 1996.
The plot centres on a young English orphan boy who enters a gigantic, magical peach, and has a wild and surreal cross-world adventure with seven magically-altered garden bugs he meets. Roald Dahl was originally going to write about a giant cherry, but changed it to James and the Giant Peach because a peach is "prettier, bigger and squishier than a cherry."Because of the story's occasional macabre and potentially frightening content, it has become a regular target of censors. The actors Jeremy Irons, Andrew Sachs, and Julian Rhind-Tutt provide the English language audiobook recordings.Rebbie Jackson
Maureen Reillette "Rebbie" Jackson-Brown (; born May 29, 1950) is an American singer. Born and raised in Gary, Indiana, she is the eldest child of the Jackson family of musicians. She first performed on stage with her siblings during shows in Las Vegas, Nevada, at the MGM Grand Hotel and Casino in 1974, before subsequently appearing in the television series The Jacksons. Her sister La Toya was born on her sixth birthday. At age 34, Jackson released her debut album Centipede (1984). The album featured songs written by Smokey Robinson, Prince, and Jackson's younger brother Michael, whose contribution (the title track "Centipede") became Rebbie's most successful single release. By the end of the 1980s, the singer had released two more albums in quick succession: Reaction (1986) and R U Tuff Enuff (1988).
Following a 10-year hiatus in her musical career, Jackson returned with the 1998 album Yours Faithfully. The production of the album, her last to date, was a collaboration with artists and producers such as Men of Vizion's Spanky Williams, Keith Thomas, and Eliot Kennedy. It also featured contributions from her children. In 2011, Rebbie embarked on the Pick Up the Phone Tour, which is dedicated to teens from all over the U.S. who have committed suicide.Scolopendra gigantea
Scolopendra gigantea, also known as the Peruvian giant yellow-leg centipede or Amazonian giant centipede, is one of the largest centipedes of the genus Scolopendra with a length up to 30 centimetres (12 in). This species is found in various places in South America and the Caribbean, where it preys on a wide variety of animals, including other sizable arthropods, amphibians, mammals and reptiles.Scutigera coleoptrata
Scutigera coleoptrata is a small, typically yellowish-grey centipede with up to 15 pairs of long legs. Originating in the Mediterranean region, the species has spread to other parts of the world, where it can live in human homes, thus gaining the name house centipede. It is an insectivore; it kills and eats other arthropods, such as insects and arachnids.Sea serpent
A sea serpent or sea dragon is a type of dragon described in various mythologies, most notably Greek (Cetus, Echidna, Hydra, Scylla), Mesopotamian (Tiamat), Hebrew (Leviathan), and Norse (Jörmungandr).Tender (rail)
A tender or coal-car is a special rail vehicle hauled by a steam locomotive containing its fuel (wood, coal, or oil) and water. Steam locomotives consume large quantities of water compared to the quantity of fuel, so their tenders are necessary to keep them running over long distances. A locomotive that pulls a tender is called a tender locomotive. Locomotives that do not have tenders and carry all their fuel and water on board the locomotive (itself) instead are called tank locomotives.
A brake tender is a tender that is heavy and used (primarily) to provide greater braking efficiency.The Human Centipede (First Sequence)
The Human Centipede (First Sequence) is a 2009 Dutch horror film written, directed, and co-produced by Tom Six. The film tells the story of a German surgeon who kidnaps three tourists and joins them surgically, mouth to anus, forming a "human centipede", a conjoined triplet. It stars Dieter Laser as the creator of the centipede, Dr. Josef Heiter, with Ashley C. Williams, Ashlynn Yennie, and Akihiro Kitamura as his victims. According to Six, the concept for the film arose from a joke he made with friends about punishing a child molester by stitching his mouth to the anus of a "fat truck driver". Inspiration for the film also came from Nazi medical experiments carried out during World War II, such as the crimes of Josef Mengele at the Auschwitz concentration camp.
When approaching investors prior to filming, Six did not mention the mouth-to-anus aspect of the plot, fearing it would put off potential backers. The financiers of The Human Centipede did not discover the full nature of the film until it was complete. The film received generally mixed reviews from film critics, but it won several accolades at international film festivals. The film received a limited theatrical release in the United States on 30 April 2010. Two sequels, titled Full Sequence and Final Sequence, also written and directed by Six, were released in 2011 and 2015, respectively. The entire trilogy was compiled into a single film in 2016, titled Complete Sequence, which Six described as a "movie centipede" due to each Sequence leading into its successor, while simultaneously working as a separate standalone film.The Human Centipede 2 (Full Sequence)
The Human Centipede 2 (Full Sequence) is a 2011 British-Dutch black and white exploitation-horror film written, directed, and co-produced by Dutch filmmaker Tom Six. The sequel to Six's 2009 film The Human Centipede (First Sequence), the film stars Laurence R. Harvey as a mentally impaired English man who watches and becomes obsessed with the first Human Centipede film, and decides to make his own "centipede" consisting of 12 people, including Ashlynn Yennie, an actress from the first film.
The film was subject to heavy censorship throughout the world, where it was sometimes edited to remove objectionable content or banned altogether. It received generally negative reviews, with much criticism focused on its acting, plot and violence.The Human Centipede 3 (Final Sequence)
The Human Centipede 3 (Final Sequence) is a 2015 English-language Dutch satirical body horror film, written and directed by Tom Six. It is the third and final installment of Six's The Human Centipede trilogy.
Starring Dieter Laser and Laurence R. Harvey, the leading actors from the first two films, in new roles, The Human Centipede 3 was released both theatrically and on video on demand on 22 May 2015. The film received predominantly negative reviews from critics.Tom Six
Tom Six (born 29 August 1973) is a Dutch filmmaker, writer, director, actor, and producer. He is best known for his trilogy of body horror films, The Human Centipede (First Sequence), The Human Centipede 2 (Full Sequence) and The Human Centipede 3 (Final Sequence).Woodlouse
A woodlouse (plural woodlice) is a crustacean from the monophyletic suborder Oniscidea within the isopods. The first woodlice were marine isopods which are presumed to have colonised land in the Carboniferous. They have many common names and although often referred to as 'terrestrial Isopods' some species live semiterrestrially or have recolonised aquatic environments. Woodlice in the families Armadillidae, Armadillidiidae, Eubelidae, Tylidae and some other genera can roll up into an almost perfect sphere as a defensive mechanism, others have partial rolling ability but most cannot conglobate at all.
Woodlice have a basic morphology of a segmented, dorso-ventrally flattened body with 7 pairs of jointed legs, specialised appendages for respiration and like other peracarids, females carry fertilised eggs in their marsupium, through which they provide developing embryos with water, oxygen and nutrients. The immature young hatch as mancae and receive further maternal care in some species. Juveniles then go through a series of moults before reaching maturity.
While the broader phylogeny of the Oniscideans has not been settled, five Infraorders/Sections are agreed on with 3637 species validated in scientific literature in 2004 and 3710 species in 2014 out of an estimated total of 5000-7000 species extant worldwide. Key adaptations to terrestrial life has led to a highly diverse set of animals; from the marine littoral zone and subterranean lakes to arid deserts and mountain slopes 4,725m above sea-level, woodlice have established themselves in most terrestrial biomes and represent the full range of transitional forms and behaviours for living on land.
Woodlice are widely studied in the contexts of evolutionary biology, behavioural ecology and nutrient cycling. They are popular as terrarium pets because of their varied colour and texture forms, conglobating ability and ease of care.Ōmukade
Ōmukade (おおむかで, "giant centipede") is a Yōkai in Japanese mythology.
Extant Arthropoda classes by subphylum