The Celyphidae, commonly known as beetle flies or beetle-backed flies, are a family of flies (order Diptera). About 115 species in about 9 genera are known chiefly from the Oriental and Afrotropic biogeographic regions with one lineage in the New World.
Celyphidae are small to medium-sized and easily recognised. The scutellum is enlarged, and forming a protective shell over the abdomen, giving them a beetle-like appearance. Also, like many beetles, Celyphidae are often shiny or metallic in color. The head has few bristles. The wings, when at rest, are folded beneath the scutellar "shell". The arista of the antenna is often flattened and leaf-like at the base. The family name is derived from the Greek word κέλνφος for pod or shell. Male celyphids lack an aedeagus and instead have paired gonapophyses that are used in copulation and are of key taxonomic value.
The biology of the family is poorly known. Adults are found along streams and rivers, and in wet, grassy areas. Larvae are saprophagous.
The family Celyphidae is considered by most authors to be the sister taxon of the Lauxaniidae (e.g., Griffiths 1972), and this has been supported by some molecular studies which suggest the (Chamaemyiidae + (Lauxaniidae + Celyphidae)) within the Lauxanoiodea. In the past they have occasionally been considered a specialized lineage within the Lauxaniidae.
Data related to Celyphidae at Wikispecies
The Acalyptratae or Acalyptrata are a subsection of the Schizophora, which are a section of the order Diptera, the "true flies". In various contexts the Acalyptratae also are referred to informally as the acalyptrate muscoids, or acalyptrates, as opposed to the Calyptratae. All forms of the name refer to the lack of calypters in the members of this subsection of flies. An alternative name, Acalypterae is current, though in minority usage. It was first used by Justin Pierre Marie Macquart in 1835 for a section of his tribe Muscides; he used it to refer to all acalyptrates plus scathophagids and phorids, but excluding Conopidae.
The confusing forms of the names stem from their first usage; Acalyptratae and Acalyptrata actually are adjectival forms in New Latin. They were coined in the mid 19th century in contexts such as "Muscae Calyptratae and Acalyptratae" and "Diptera Acalyptrata", and the forms stuck.The Acalyptratae are a large assemblage, exhibiting very diverse habits, with one notable and perhaps surprising exception: no known acalyptrates are obligate blood-feeders (hematophagous), though blood feeding at various stages of the life history is common throughout other Dipteran sections.Acelyphus
Acelyphus is a genus of beetle flies. It is known from the Oriental ecozone.Afrocelyphus
Afrocelyphus is a genus of Chloropidae, the genus was originally placed in the family Celyphidae.Asilomorpha
The Brachyceran infraorder Asilomorpha is a large and diverse group of flies, containing the bulk of the nonmuscoid Brachycera. The larvae of asilomorphs are extremely diverse in habits, as well.Calyptratae
Calyptratae is a subsection of Schizophora in the insect order Diptera, commonly referred to as the calyptrate muscoids (or simply calyptrates). It consists of those flies which possess a calypter that covers the halteres, among which are some of the most familiar of all flies, such as the house fly.
About 18,000 described species are in this group, or about 12% of all the flies yet described.Carnoidea
Carnoidea are a superfamily of Acalyptratae flies.Celyphus
Celyphus is a genus of beetle flies. It is known from the Oriental and African ecozones. Up to 1859, all species of beetle flies were placed in this genus.Chamaecelyphus
Chamaecelyphus is a genus of Celyphidae.Chironomoidea
The Chironomoidea are a superfamily within the order Diptera, suborder Nematocera, infraorder Culicomorpha. This superfamily contains the families Chironomidae, Ceratopogonidae, Simuliidae, and Thaumaleidae. One of the more important characteristics used to define them is the form of the larval mouthparts.Culicoidea
The Culicoidea are a superfamily within the order Diptera. The following families are included within the Culicoidea:
Dixidae – meniscus midges
Corethrellidae – frog-biting midges
Chaoboridae – phantom midges
Culicidae – mosquitoesCulicomorpha
The Culicomorpha are an infraorder of Nematocera, including mosquitoes, black flies, and several extant and extinct families of insects.Idiocelyphus
Idiocelyphus is a genus of beetle flies. It is known from the Oriental ecozone. Most are from the Philippines.Lauxanioidea
The Lauxanioidea are a superfamily of flies that includes the two large families, the Lauxaniidae and Chamaemyiidae, and the small family Celyphidae. Generally, they are small to medium, densely populated, coloured flies. The Chamaemyiidae and Cremifaniidae live as parasites on insects. The family Celyphidae look like beetles.
Some authors also recognize the family Cremifaniidae, but most place this in the Chamaemyiidae.Oestroidea
Oestroidea is a superfamily of Calyptratae including the blow flies, bot flies, flesh flies, and their relatives.The superfamily includes the families:
Mesembrinellidae (formerly included in Calliphoridae)
Rhiniidae (formerly included in Calliphoridae)
Sciaroidea is a superfamily in the infraorder Bibionomorpha. There are about 16 families and more than 15,000 described species in Sciaroidea. Most of its constituent families are various gnats.Spaniocelyphus
Spaniocelyphus is a genus of beetle flies known from the Oriental ecozone.Tabanoidea
Superfamily Tabanoidea are insects in the order Diptera.Tephritoidea
The Tephritoidea are a superfamily of flies. The following families are included:
Pallopteridae — flutter flies
Piophilidae — skippers
Platystomatidae — signal flies
Tephritidae — fruit flies
Ulidiidae (Otitidae) — picture-winged fliesThe Tachiniscinae, formerly ranked as the family Tachiniscidae, are now included in the Tephritidae.Tipulomorpha
The Tipulomorpha are an infraorder of Nematocera, containing the crane flies, a very large group, and allied families.
One recent classification based largely on fossils splits this group into a series of extinct superfamilies (below), and includes members of other infraorders, but this has not gained wide acceptance.
Extant Diptera families