CITES (the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals. It was drafted as a result of a resolution adopted in 1963 at a meeting of members of the International Union for Conservation of Nature (IUCN). The convention was opened for signature in 1973 and CITES entered into force on 1 July 1975. Its aim is to ensure that international trade in specimens of wild animals and plants does not threaten the survival of the species in the wild, and it accords varying degrees of protection to more than 35,000 species of animals and plants. In order to ensure that the General Agreement on Tariffs and Trade (GATT) was not violated, the Secretariat of GATT was consulted during the drafting process.[1]

As of 2018, Secretary-General of the CITES Secretariat is Ivonne Higuero.[2]

The Convention on International Trade in Endangered Species
Logo of the 40th anniversary of CITES
Signed3 March 1973
LocationUnited States Washington, D.C.
Effective1 July 1975
Condition10 ratifications
DepositarySwitzerland Government of the Swiss Confederation
Convention on International Trade in Endangered Species of Wild Fauna and Flora at Wikisource

Background and operation

CITES is one of the largest and oldest conservation and sustainable use agreements in existence. Participation is voluntary, and countries that have agreed to be bound by the Convention are known as Parties. Although CITES is legally binding on the Parties, it does not take the place of national laws. Rather it provides a framework respected by each Party, which must adopt their own domestic legislation to implement CITES at the national level. Often, domestic legislation is either non-existent (especially in Parties that have not ratified it), or with penalties with the gravity of the crime and insufficient deterrents to wildlife traders.[3] As of 2002, 50% of Parties lacked one or more of the four major requirements for a Party: designation of Management and Scientific Authorities; laws prohibiting the trade in violation of CITES; penalties for such trade; laws providing for the confiscation of specimens.[4]

Funding for the activities of the Secretariat and Conference of the Parties (CoP) meetings comes from a Trust Fund derived from Party contributions. Trust Fund money is not available to Parties to improve implementation or compliance. These activities, and all those outside Secretariat activities (training, species specific programmes such as Monitoring the Illegal Killing of Elephants - MIKE) must find external funding, mostly from donor countries and regional organizations such as the European Union.

Although the Convention itself does not provide for arbitration or dispute in the case of noncompliance, 36 years of CITES in practice has resulted in several strategies to deal with infractions by Parties. The Secretariat, when informed of an infraction by a Party, will notify all other parties. The Secretariat will give the Party time to respond to the allegations and may provide technical assistance to prevent further infractions. Other actions the Convention itself does not provide for but that derive from subsequent COP resolutions may be taken against the offending Party. These include:

  • Mandatory confirmation of all permits by the Secretariat
  • Suspension of cooperation from the Secretariat
  • A formal warning
  • A visit by the Secretariat to verify capacity
  • Recommendations to all Parties to suspend CITES related trade with the offending party[5]
  • Dictation of corrective measures to be taken by the offending Party before the Secretariat will resume cooperation or recommend resumption of trade

Bilateral sanctions have been imposed on the basis of national legislation (e.g. the USA used certification under the Pelly Amendment to get Japan to revoke its reservation to hawksbill turtle products in 1991, thus reducing the volume of its exports).

Infractions may include negligence with respect to permit issuing, excessive trade, lax enforcement, and failing to produce annual reports (the most common).

Originally, CITES addressed depletion resulting from demand for luxury goods such as furs in Western countries, but with the rising wealth of Asia, particularly in China, the focus changed to products demanded there, particularly those used for luxury goods such as ivory or shark fins or for superstitious purposes such as rhinoceros horn. As of 2013 the demand was massive and had expanded to include thousands of species previously considered unremarkable and in no danger of extinction such as manta rays or pangolins.[6]


Parties to the treaty. Greenland is covered by CITES regulations through Denmark.[7]

The text of the Convention was finalized at a meeting of representatives of 80 countries in Washington, D.C., United States, on 3 March 1973. It was then open for signature until 31 December 1974. It entered into force after the 10th ratification by a signatory country, on 1 July 1975. Countries that signed the Convention become Parties by ratifying, accepting or approving it. By the end of 2003, all signatory countries had become Parties. States that were not signatories may become Parties by acceding to the Convention. As of October 2016, the Convention has 183 parties, including 182 states and the European Union.[8]

The CITES Convention includes provisions and rules for trade with non-Parties. All member states of the United Nations are party to the treaty, with the exception of Andorra, Democratic People's Republic of Korea, Federated States of Micronesia, Haiti, Kiribati, Marshall Islands, Nauru, South Sudan, East Timor, Tonga, Turkmenistan, and Tuvalu. UN observer the Holy See is also not a member. The Faroe Islands, an autonomous country in the Kingdom of Denmark, is also treated as a non-Party to CITES (both the Danish mainland and Greenland are part of CITES).[7][9]

An amendment to the text of the Convention, known as the Gaborone Amendment[10] allows regional economic integration organizations (REIO), such as the European Union, to have the status of a member state and to be a Party to the Convention. The REIO can vote at CITES meetings with the number of votes representing the number of members in the REIO, but it does not have an additional vote.

In accordance with Article XVII, paragraph 3, of the CITES Convention, the Gaborone Amendment entered into force on 29 November 2013, 60 days after 54 (two-thirds) of the 80 States that were party to CITES on 30 April 1983 deposited their instrument of acceptance of the amendment. At that time it entered into force only for those States that had accepted the amendment. The amended text of the Convention will apply automatically to any State that becomes a Party after 29 November 2013. For States that became party to the Convention before that date and have not accepted the amendment, it will enter into force 60 days after they accept it.[11]

Regulation of trade

CITES works by subjecting international trade in specimens of selected species to certain controls. All import, export, re-export and introduction from the sea of species covered by the Convention has to be authorized through a licensing system. According to Article IX of the Convention, Management and Scientific Authorities, each Party to the Convention must designate one or more Management Authorities in charge of administering that licensing system and one or more Scientific Authorities to advise them on the effects of trade on the status of CITES-listed species.


Roughly 5,000 species of animals and 29,000 species of plants are protected by CITES against over-exploitation through international trade. Each protected species or population is included in one of three lists, called appendices[12][13] (explained below). The Appendix that lists a species or population reflects the extent of the threat to it and the controls that apply to the trade.

Species may be split-listed meaning that some populations of a species are on one Appendix, while some are on another. Some people argue that this is risky as specimens from a more protected population could be 'laundered' through the borders of a Party whose population is not as strictly protected. The African bush elephant (Loxodonta africana) is currently split-listed, with all populations except those of Botswana, Namibia, South Africa and Zimbabwe listed in Appendix I. Those of Botswana, Namibia, South Africa and Zimbabwe are listed in Appendix II. Listing the species over the whole of its range would prevent such 'laundering' but also restricts trade in wildlife products by range states with good management practices. There are also species that have only some populations listed in an Appendix. One example is the pronghorn (Antilocapra americana), a ruminant native to North America. Its Mexican population is listed in Appendix I, but its U.S. and Canadian populations are not listed (though certain U.S. populations in Arizona are nonetheless protected under the Endangered Species Act).

Species are proposed for inclusion in or deletion from the Appendices at meetings of the Conference of the Parties (CoP), which are held approximately once every three years, the most recent of which was CoP (CoP 17) in Johannesburg, South Africa from 24 September to 5 October 2016 at the Sandton Convention Center.[14]

Species in the Appendices may be proposed for addition, change of Appendix, or de-listing (i.e., deletion) by any Party, whether or not it is a range State and changes may be made despite objections by range States if there is sufficient (2/3 majority) support for the listing. These discussions are usually among the most contentious at CoP meetings.

There has been increasing willingness within the Parties to allow for trade in products from well-managed populations. For instance, sales of the South African white rhino have generated revenues that helped pay for protection. Listing the species on Appendix I increased the price of rhino horn (which fueled more poaching), but the species survived wherever there was adequate on-the-ground protection. Thus field protection may be the primary mechanism that saved the population, but it is likely that field protection would not have been increased without CITES protection.[15]

Appendix I

Appendix I, about 1200 species, are species that are threatened with extinction and are or may be affected by trade. Commercial trade in wild-caught specimens of these species is illegal (permitted only in exceptional licensed circumstances). Captive-bred animals or cultivated plants of Appendix I species are considered Appendix II specimens, with concomitant requirements (see below and Article VII). The Scientific Authority of the exporting country must make a non-detriment finding, assuring that export of the individuals will not adversely affect the wild population. Any trade in these species requires export and import permits. The Management Authority of the exporting state is expected to check that an import permit has been secured and that the importing state is able to care for the specimen adequately. Notable animal species listed in Appendix I include the red panda (Ailurus fulgens), western gorilla (Gorilla gorilla), the chimpanzee species (Pan spp.), tigers (Panthera tigris subspecies), Asiatic lion (Panthera leo persica), leopards (Panthera pardus), jaguar (Panthera onca), cheetah (Acinonyx jubatus), Asian elephant (Elephas maximus), some populations of African bush elephant (Loxodonta africana), the dugong and manatees (Sirenia), and all rhinoceros species (except some Southern African subspecies populations).[16]

Appendix II

Appendix II, about 21,000 species, are species that are not necessarily threatened with extinction, but may become so unless trade in specimens of such species is subject to strict regulation in order to avoid utilization incompatible with the survival of the species in the wild. In addition, Appendix II can include species similar in appearance to species already listed in the Appendices. International trade in specimens of Appendix II species may be authorized by the granting of an export permit or re-export certificate. In practice, many hundreds of thousands of Appendix II animals are traded annually.[17] No import permit is necessary for these species under CITES, although some Parties do require import permits as part of their stricter domestic measures. A non-detriment finding and export permit are required by the exporting Party.[16]

In addition, Article VII of CITES states that specimens of animals listed in Appendix I that are bred in captivity for commercial purposes are treated as Appendix II. The same applies for specimens of Appendix I plants artificially propagated for commercial purposes.[18]

Examples of species listed on Appendix II are the great white shark (Carcharodon carcharias), the American black bear (Ursus americanus), Hartmann's mountain zebra (Equus zebra hartmannae), green iguana (Iguana iguana), queen conch (Strombus gigas), Emperor scorpion (Pandinus imperator), Mertens' water monitor (Varanus mertensi), bigleaf mahogany (Swietenia macrophylla) and lignum vitae "ironwood" (Guaiacum officinale).

Appendix III

Appendix III, about 170 species, are species that are listed after one member country has asked other CITES Parties for assistance in controlling trade in a species. The species are not necessarily threatened with extinction globally. In all member countries, trade in these species is only permitted with an appropriate export permit and a certificate of origin from the state of the member country who has listed the species.[16]

Examples of species listed on Appendix III and the countries that listed them are the two-toed sloth (Choloepus hoffmanni) by Costa Rica, sitatunga (Tragelaphus Spekii) by Ghana, African civet (Civettictis civetta) by Botswana, and alligator snapping turtle by the USA.

Amendments and reservations

Amendments to the Convention must be supported by a two-thirds majority who are "present and voting" and can be made during an extraordinary meeting of the COP if one-third of the Parties are interested in such a meeting. The Gaborone Amendment (1983) allows regional economic blocs to accede to the treaty. Reservations (Article XXIII) can be made by any Party with respect to any species, which considerably weakens the treaty (see [1] for current reservations). Trade with non-Party states is allowed, although permits and certificates are recommended to be issued by exporters and sought by importers.

Notable reservations include those by Iceland, Japan and Norway on various baleen whale species and those on Falconiformes by Saudi Arabia.[19]

Shortcomings and concerns

Approach to biodiversity conservation

General limitations about the structure and philosophy of CITES include: by design and intent it focuses on trade at the species level and does not address habitat loss, ecosystem approaches to conservation, or poverty; it seeks to prevent unsustainable use rather than promote sustainable use (which generally conflicts with the Convention on Biological Diversity), although this has been changing (see Nile crocodile, African elephant, South African white rhino case studies in Hutton and Dickinson 2000). It does not explicitly address market demand.[20] Funding does not provide for increased on-the-ground enforcement (it must apply for bilateral aid for most projects of this nature).


By design, CITES regulates and monitors trade in the manner of a "negative list" such that trade in all species is permitted and unregulated unless the species in question appears on the Appendices or looks very much like one of those taxa. Then and only then, trade is regulated or constrained. Because the remit of the Convention covers millions of species of plants and animals, and tens of thousands of these taxa are potentially of economic value, in practice this negative list approach effectively forces CITES signatories to expend limited resources on just a select few, leaving many species to be traded with neither constraint nor review. For example, recently several bird classified as threatened with extinction appeared in the legal wild bird trade because the CITES process never considered their status. If a "positive list" approach were taken, only species evaluated and approved for the positive list would be permitted in trade, thus lightening the review burden for member states and the Secretariat, and also preventing inadvertent legal trade threats to poorly known species.

Specific weaknesses in the text include: it does not stipulate guidelines for the 'non-detriment' finding required of national Scientific Authorities; non-detriment findings require copious amounts of information; the 'household effects' clause is often not rigid enough/specific enough to prevent CITES violations by means of this Article (VII); non-reporting from Parties means Secretariat monitoring is incomplete; and it has no capacity to address domestic trade in listed species.

Reform suggestions

Suggestions for improvement in the operation of CITES include: more regular missions by the Secretariat (not reserved just for high-profile species); improvement of national legislation and enforcement; better reporting by Parties (and the consolidation of information from all sources-NGOs, TRAFFIC, the wildlife trade monitoring network and Parties); more emphasis on enforcement-including a technical committee enforcement officer; the development of CITES Action Plans (akin to Biodiversity Action Plans related to the Convention on Biological Diversity) including: designation of Scientific/Management Authorities and national enforcement strategies; incentives for reporting and timelines for both Action Plans and reporting. CITES would benefit from access to Global Environment Facility (GEF), funds-although this is difficult given the GEFs more ecosystem approach-or other more regular funds. Development of a future mechanism similar to that of the Montreal Protocol (developed nations contribute to a fund for developing nations) could allow more funds for non-Secretariat activities.[4]

On 15 July 2008, the Committee of Environmental Insecticides that oversees the administration of the convention between meetings of all the Parties granted China and Japan permission to import elephant ivory from four African government stockpiles, the ivory being sold at a single auction in each country. The amounts to be sold comprise approximately 44 tons from Botswana, 9 tons from Namibia, 51 tons from South Africa, and 4 tons from Zimbabwe. The Chinese government in 2003 acknowledged that it had lost track of 121 tons of ivory between 1991 and 2002.


From 2005 – 2009 the legal trade corresponded with these numbers

  • 317,000 live birds
  • More than 2 million live reptiles
  • 2.5 million crocodile skins
  • 2.1 million snake skins
  • 73 tons of caviar
  • 1.1 million beaver skins
  • Millions of pieces of coral
  • 20,000 mammalian hunting trophies

In the 1990s the annual trade of legal animal products was $160 billion annually. In 2009 the estimated value almost doubled to $300 billion.[21]


The Conference of the Parties (CoP) is held once every three years. The last Conference of the Parties (CoP 17) was held in Johannesburg, South Africa, and the one before it (CoP 16) was held in Bangkok, Thailand, in 2013. The next one (CoP 18) will be in Sri Lanka in 2019. The location of the next CoP is chosen at the close of each CoP by a secret ballot vote.

The CITES Committees (Animals Committee, Plants Committee and Standing Committee) hold meetings during each year that does not have a CoP, while the Standing committee meets also in years with a CoP. The Committee meetings take place in Geneva, Switzerland (where the Secretariat of the CITES Convention is located), unless another country offers to host the meeting. The Secretariat is administered by UNEP. The Animals and Plants Committees have sometimes held joint meetings. The previous joint meeting was held in March 2012 in Dublin, Ireland, and the latest one was held in Veracruz, Mexico in May 2014.

Meeting City Country Duration
CoP 1 Bern   Switzerland 2–6 November 1976
CoP 2 San José  Costa Rica 19–30 March 1979
CoP 3 New Delhi  India 25 February – 8 March 1981
CoP 4 Gaborone  Botswana 19 – 30 April 1983
CoP 5 Buenos Aires  Argentina 22 April – 3 May 1985
CoP 6 Ottawa  Canada 12–24 July 1987
CoP 7 Lausanne   Switzerland 9–20 October 1989
CoP 8 Kyoto  Japan 2–13 March 1992
CoP 9 Fort Lauderdale  United States 7–18 November 1994
CoP 10 Harare  Zimbabwe 9–20 June 1997
CoP 11 Gigiri  Kenya 10–20 April 2000
CoP 12 Santiago  Chile 3–15 November 2002
CoP 13 Bangkok  Thailand 2–14 October 2004
CoP 14 The Hague  Netherlands 3–15 June 2007
CoP 15 Doha  Qatar 13–25 March 2010
CoP 16 Bangkok  Thailand 3–14 March 2013
CoP 17 Johannesburg  South Africa 24 September – 5 October 2016
CoP 18 Colombo  Sri Lanka 2019

A current list of upcoming meetings appears on the CITES calendar at

See also


  1. ^ "What is CITES?". CITES. Retrieved 13 February 2012.
  2. ^ "Ivonne Higuero named as new CITES Secretary-General". CITES. Retrieved 3 December 2018.
  3. ^ "Zimmerman, "The Black Market for Wildlife: llegal Wildlife Trade," Vanderbilt Journal of Transnational Law 36 (5) 1657-1689 (November 2003)". Archived from the original on 21 June 2010. Retrieved 23 March 2010.
  4. ^ a b Reeve, Policing International Trade in Endangered Species: The CITES Treaty and agreement. London: Earthscan, 2000.
  5. ^ "Countries currently subject to a recommendation to suspend trade". CITES. Retrieved 13 February 2012.
  6. ^ Bettina Wassener (12 March 2013). "No Species Is Safe From Burgeoning Wildlife Trade". The New York Times. Retrieved 13 March 2013.
  7. ^ a b CITES: Denmark. Retrieved 24 February 2015.
  8. ^ "List of Contracting Parties - CITES".
  9. ^ CITES: Faroe Islands. Retrieved 24 February 2015.
  10. ^ "Gaborone amendment to the text of the Convention". CITES. Retrieved 13 February 2012.
  11. ^ "Gaborone amendment to the text of the Convention - CITES".
  12. ^ Convention on International Trade in Endangered Species of Wild Fauna and Flora (2013). "Appendices I, II and III". Retrieved 13 September 2013.
  13. ^ Convention on International Trade in Endangered Species of Wild Fauna and Flora. "The CITES Appendices". Retrieved 13 September 2013.
  14. ^ "CITES Calendar". CITES. Retrieved 13 February 2012.
  15. ^ Hutton and Dickinson, Endangered Species Threatened Convention: The Past, Present and Future of CITES. London: Africa Resources Trust, 2000.
  16. ^ a b c "Appendices I, II and III". CITES. Retrieved 13 February 2012.
  17. ^ "CITES Export Quotas". CITES. Retrieved 13 February 2012.
  18. ^ "Article VII: Exemptions and Other Special Provisions Relating to Trade". Convention on International Trade in Endangered Species of Wild Fauna and Flora. CITES. Retrieved 17 October 2014.
  19. ^ "CITES Reservations Entered by Parties". CITES. Retrieved 13 February 2012.
  20. ^ Hill, 1990, "The Convention on International Trade in Endangered Species: Fifteen Years Later," Loyola of Los Angeles International and Comparative Law Journal 13: 231
  21. ^ Ceballos, G.; Ehrlich, A. H.; Ehrlich, P. R. (2015). The Annihilation of Nature: Human Extinction of Birds and Mammals. Baltimore, Maryland: Johns Hopkins University Press. pp. 150. ISBN 1421417189 – via Open Edition.

Further reading

External links

Member countries (Parties)
Lists of species included in Appendices I, II and III (i.e. species protected by CITES)

The barbiton, or barbitos (Gr: βάρβιτον or βάρβιτος; Lat. barbitus), is an ancient stringed instrument known from Greek and Roman classics related to the lyre. The barbat or barbud, also sometimes called barbiton, is an unrelated lute-like instrument derived from Persia.

The Greek instrument was a bass version of the kithara, and belonged in the zither family, but in medieval times, the same name was used to refer to a different instrument that was a variety of lute.


Cocobolo is a tropical hardwood of Central American trees belonging to the genus Dalbergia. Only the heartwood of cocobolo is used; it is usually orange or reddish-brown, often with darker irregular traces weaving through the wood. The heartwood changes color after being cut, and can be polished to a lustrous, glassy finish; being quite dense, some has a specific gravity of over 1.0, and will sink in water. The sapwood (not often used) is a creamy yellow, with a sharp boundary between it and the heartwood.

Comedy horror

Comedy horror is a literary and film genre that combines elements of comedy and horror fiction. Comedy horror has been described as able to be categorized under three types: "black comedy, parody and spoof." It often crosses over with the black comedy genre. Comedy horror can also parody or subtly spoof horror clichés as its main source of humour or use those elements to take a story in a different direction, for example in The Cabin in the Woods.

Author Bruce G. Hallenbeck cites the short story "The Legend of Sleepy Hollow" by Washington Irving as "the first great comedy horror story". The story made readers "laugh one moment and scream the next", and its premise was based on mischief typically found during the holiday Halloween.

Conservation status

The conservation status of a group of organisms (for instance, a species) indicates whether the group still exists and how likely the group is to become extinct in the near future. Many factors are taken into account when assessing conservation status: not simply the number of individuals remaining, but the overall increase or decrease in the population over time, breeding success rates, and known threats. Various systems of conservation status exist and are in use at international, multi-country, national and local levels as well as for consumer use.

Edward the Black Prince

Edward of Woodstock, known to history as the Black Prince (15 June 1330 – 8 June 1376), was the eldest son of King Edward III of England, and thus the heir to the English throne. He died before his father and so his son, Richard II, succeeded to the throne instead. Edward nevertheless still earned distinction as one of the most successful English commanders during the Hundred Years' War, being regarded by his contemporaries as a model of chivalry and one of the greatest knights of his age.Edward was created Duke of Cornwall in 1337. He was guardian of the kingdom in his father's absence in 1338, 1340, and 1342. He was created Prince of Wales in 1343 and knighted by his father at La Hogne in 1346.

In 1346 Prince Edward commanded the vanguard at the Battle of Crécy, his father intentionally leaving him to win the battle. He took part in Edward III's 1349 Calais expedition. In 1355 he was appointed the king's lieutenant in Gascony, and ordered to lead an army into Aquitaine on a chevauchée, during which he pillaged Avignonet and Castelnaudary, sacked Carcassonne, and plundered Narbonne. The next year (1356) on another chevauchée he ravaged Auvergne, Limousin, and Berry but failed to take Bourges. He offered terms of peace to King John II of France, who had outflanked him near Poitiers, but refused to surrender himself as the price of their acceptance. This led to the Battle of Poitiers where his army routed the French and took King John prisoner.

The year after Poitiers, the Black Prince returned to England. In 1360 he negotiated the treaty of Bretigny. He was created Prince of Aquitaine and Gascony in 1362, but his suzerainty was not recognised by the lord of Albret or other Gascon nobles. He was directed by his father to forbid the marauding raids of the English and Gascon free companies in 1364. He entered into an agreement with don Pedro of Castile and Charles II of Navarre, by which Pedro covenanted to mortgage Castro de Urdiales and the province of Biscay to him as security for a loan; in 1366 a passage was thus secured through Navarre. In 1367 he received a letter of defiance from Henry of Trastámara, Don Pedro's half-brother and rival. The same year, after an obstinate conflict, he defeated Henry at the Battle of Nájera. However, after a wait of several months, during which he failed to obtain either the province of Biscay or liquidation of the debt from Don Pedro, he returned to Aquitaine. Prince Edward persuaded the estates of Aquitaine to allow him a hearth tax of ten sous for five years in 1368, thereby alienating the lord of Albret and other nobles. Drawn into open war with Charles V of France in 1369, he took Limoges, where in 1370 he gave orders for an indiscriminate massacre in revenge for the voluntary surrender of that town to the French by its bishop, who had been his private friend.

The Black Prince returned to England in 1371 and the next year resigned the principality of Aquitaine and Gascony. He led the commons in their attack upon the Lancastrian administration in 1376. He died in 1376 of dysentery and was buried in Canterbury Cathedral, where his surcoat, helmet, shield, and gauntlets are still preserved.

Hanes–Woolf plot

In biochemistry, a Hanes–Woolf plot is a graphical representation of enzyme kinetics in which the ratio of the initial substrate concentration [S] to the reaction velocity v is plotted against [S]. It is based on the rearrangement of the Michaelis–Menten equation shown below:

where Km is the Michaelis–Menten constant and Vmax is the maximum reaction velocity.

J B S Haldane stated that this method was due to Barnet Woolf. It was also used by Charles Samuel Hanes, even though he neither mentions nor cites Woolf. Hanes pointed out that the use of linear regression to determine kinetic parameters from this type of linear transformation is flawed, because it generates the best fit between observed and calculated values of 1/v, rather than v.

The equation can be derived from the Michaelis–Menten equation as follows:

invert and multiply by [S]:


As is clear from the equation, perfect data will yield a straight line of slope 1/Vmax, a y-intercept of Km/Vmax and an x-intercept of −Km.

Like other techniques that linearize the Michaelis–Menten equation, the Hanes–Woolf plot was used historically for rapid determination of the important kinetic parameters Km, Vmax and Vmax/Km, but it has been superseded by nonlinear regression methods that are significantly more accurate and no longer computationally inaccessible. It remains useful, however, as a means to present data graphically.

One drawback of the Hanes–Woolf approach is that neither ordinate nor abscissa represent independent variables: both are dependent on substrate concentration. As a result, the typical measure of goodness of fit, the correlation coefficient R, is not applicable.


Ivory is a hard, white material from the tusks (traditionally elephants') and teeth of animals, that consists mainly of dentine, one of the physical structures of teeth and tusks. The chemical structure of the teeth and tusks of mammals is the same, regardless of the species of origin. The trade in certain teeth and tusks other than elephant is well established and widespread; therefore, "ivory" can correctly be used to describe any mammalian teeth or tusks of commercial interest which are large enough to be carved or scrimshawed. It has been valued since ancient times in art or manufacturing for making a range of items from ivory carvings to false teeth, fans, dominoes and joint tubes. Elephant ivory is the most important source, but ivory from mammoth, walrus, hippopotamus, sperm whale, killer whale, narwhal and wart hog are used as well. Elk also have two ivory teeth, which are believed to be the remnants of tusks from their ancestors.The national and international trade in ivory of threatened species such as African and Asian elephants is illegal. The word ivory ultimately derives from the ancient Egyptian âb, âbu ("elephant"), through the Latin ebor- or ebur.

Ivory trade

The ivory trade is the commercial, often illegal trade in the ivory tusks of the hippopotamus, walrus, narwhal, mammoth, and most commonly, African and Asian elephants.

Ivory has been traded for hundreds of years by people in regions such as Greenland, Alaska, and Siberia. The trade, in more recent times, has led to endangerment of species, resulting in restrictions and bans. Ivory was formerly used to make piano keys and other decorative items because of the white color it presents when processed but the piano industry abandoned ivory as a key covering material in the 1980s.

List of cities and towns in Russia by population

This is a list of cities and towns in Russia with a population of over 50,000 as of the 2010 Census. These numbers are the population within the limits of the city/town proper, not the urban area or metropolitan area figures.

The list excludes the city of Sevastopol and locations within the Republic of Crimea, since those were not subject to the 2010 Census as constituent parts of Ukraine. The city of Zelenograd (a part of the federal city of Moscow) and the municipal cities/towns of the federal city of St. Petersburg are also excluded, as they are not enumerated in the 2010 Census as stand-alone localities.

List of regencies and cities of Indonesia

This is the list of regencies and cities of Indonesia. Both regencies and cities are second-level administrative subdivision in Indonesia, immediately below the provinces, and above the districts. They are roughly equivalent to American counties.In Indonesia, both regency and city are at the same administration level, each having their own local government and legislative body. The difference between a regency and a city lies in demography, size, and economy. Generally, a regency comprises a rural area larger than a city, but also often includes various towns. A city usually has non-agricultural economic activities.

A regency (Indonesian: kabupaten) is headed by a regent, known locally as bupati, while a city (Indonesian: kota) is headed by a mayor (walikota). All regents, mayors, and members of legislatures are directly elected via elections to serve for a five-year term which can be renewed once. Each regency or city is divided further into districts more commonly known as kecamatan, or distrik in Papua.

An administrative city is a city without its own local legislatures (Dewan Perwakilan Rakyat Daerah). The mayor of an administrative city is directly appointed by the Governor. This type of city in Indonesia is only found in Jakarta which consisted of 5 administrative cities and 1 administrative regency.

Following the implementation of decentralization beginning on 1 January 2001, regencies and city municipalities became the key administrative units responsible for providing most governmental services.The list below groups regencies and cities in Indonesia by provinces. Each regency has an administrative centre, the regency seat.

Murrough O'Brien, 1st Earl of Inchiquin

Murrough MacDermod O'Brien, 6th Baron Inchiquin, 1st Baron O'Brien of Burren, 1st Earl of Inchiquin (1614 – 9 September 1674), was known as Murchadh na dTóiteán ("of the conflagrations" i.e.: extensive burnings) – of Irish who would not convert to Anglicanism and their land, crops, livestock, and dwellings.O'Brien studied war in the Spanish service and fought against the confederate Catholics on the outbreak of the Irish Rebellion of 1641. He was made governor of Munster in 1642 and had some small success, but was hampered by lack of funds. Sidney Lee states that he outwitted the Irish leader, Donough MacCarty, 2nd Viscount Muskerry, with threats and promises. O'Brien persuaded

Muskerry to delay attacking the garrisons at Cappoquin and Lismore until a truce was brokered by a representative of King Charles I, after which O'Brien forces dispersed. O'Brien visited Charles I at Oxford in 1644. He was forced to submit to parliament in 1644, as the parliamentarians being masters of the sea, were therefore the only people who could help the Munster Protestants. He was made President of Munster, supplies having been brought to him by Philip Sidney (later 3rd Earl of Leicester) in 1647. O'Brien gradually became master of the south of Ireland, and declared for Charles I in 1648, fortified the southern ports against parliament and signed a truce with the confederate Catholics. He was joined by James Butler, 1st Duke of Ormonde, with whom he got possession of Drogheda and Dundalk. O'Brien lost influence in Munster, which revolted after Cromwell's landing, 1649, but made a stand at Kilmallock in 1649. He retired to the west of the Shannon and then left Ireland for France in 1650, where he became one of the royal council and in 1654 was created Earl of Inchiquin. He served under the French in Catalonia in 1654, and was engaged in the Sexby Plot in 1656 and in the same year became a Roman Catholic. He was taken prisoner by the Algerines in 1660, but ransomed the same year and became high steward of Queen Henrietta Maria's household. He lived quietly in Ireland after 1663.

Napoleonic Wars casualties

The casualties of the Napoleonic Wars (1803–1815), direct and indirect, break down as follows:

Note that the following deaths listed include both killed in action as well as deaths from other causes; Deaths from diseases such as those from wounds; of starvation; exposure; drowning; friendly fire; and atrocities; Medical treatments were changed drastically at this time. 'Napoleon's Surgeon', Baron Dominique Jean Larrey, used horse-drawn carts as ambulances to quickly remove the wounded from the field of battle. This method became so successful that he was subsequently asked to organize the medical care for the 14 armies of the French Republic.

Parenthetical referencing

Parenthetical referencing, also known as Harvard referencing, is a citation style in which partial citations—for example, "(Smith 2010, p. 1)"—are enclosed within parentheses and embedded in the text, either within or after a sentence. They are accompanied by a full, alphabetized list of citations in an end section, usually titled "references", "reference list", "works cited", or "end-text citations". Parenthetical referencing can be used in lieu of footnote citations (the Vancouver system).

There are two styles of parenthetical referencing:

Author–date: primarily used in the natural sciences and social sciences, and recommended by the American Chemical Society and the American Psychological Association (APA);

Author–title or author–page: primarily used in the arts and the humanities, and recommended by the Modern Language Association (MLA).

Prefecture-level city

A prefectural-level municipality (Chinese: 地级市), prefectural-level city or prefectural city; formerly known as province-administrated city (Chinese: 省辖市) from 1949 to 1983, is an administrative division of the People's Republic of China (PRC), ranking below a province and above a county in China's administrative structure. Prefectural level cities form the second level of the administrative structure (alongside prefectures, leagues and autonomous prefectures). Administrative chiefs (mayors) of prefectural level cities generally have the same rank as a division chief (Chinese: 司长) of a national ministry. Since the 1980s, most former prefectures have been renamed into prefectural level cities.

A prefectural level city is a "city" (Chinese: 市; pinyin: shì) and "prefecture" (Chinese: 地区; pinyin: dìqū) that have been merged into one consolidated and unified jurisdiction. As such it is simultaneously a city, which is a municipal entry with subordinate districts, and a prefecture with subordinate county-level cities and counties which is an administrative division of a province.

A prefectural level city is often not a "city" in the usual sense of the term (i.e., a large continuous urban settlement), but instead an administrative unit comprising, typically, a main central urban area (a city in the usual sense, usually with the same name as the prefectural level city), and its much larger surrounding rural area containing many smaller cities, towns and villages. The larger prefectural level cities span over 100 kilometres (62 mi).

Prefectural level cities nearly always contain multiple counties, county level cities, and other such sub-divisions. This results from the fact that the formerly predominant prefectures, which prefectural level cities have mostly replaced, were themselves large administrative units containing cities, smaller towns, and rural areas. To distinguish a prefectural level city from its actual urban area (city in the strict sense), the term 市区 shìqū ("urban area"), is used.

The first prefectural level cities were created on 5 November 1983. Over the following two decades, prefectural level cities have come to replace the vast majority of Chinese prefectures; the process is still ongoing.

Most provinces are composed entirely or nearly entirely of prefectural level cities. Of the 22 provinces and 5 autonomous regions of the PRC, only 9 provinces (Yunnan, Guizhou, Qinghai, Heilongjiang, Sichuan, Gansu, Jilin, Hubei, Hunan) and 3 autonomous regions (Xinjiang, Tibet, Inner Mongolia) have at least one or more second level or prefectural level divisions that are not prefectural level cities.

Criteria that a prefecture must meet to become a prefectural level city:

An urban centre with a non-rural population over 250,000

gross output of value of industry of 200,000,000 RMB (US$32 million)

the output of tertiary industry supersedes that of primary industry, contributing over 35% of the GDP15 large prefectural level cities have been granted the status of sub-provincial city, which gives them much greater autonomy.

Shijiazhuang and Zhengzhou are the largest prefectural level cities with populations approaching or exceeding some sub-provincial cities.

A sub-prefecture-level city is a county-level city with powers approaching those of prefectural level cities.


A scourge is a whip or lash, especially a multi-thong type, used to inflict severe corporal punishment or self-mortification. It is usually made of leather.

The Mysterious Cities of Gold

The Mysterious Cities of Gold, originally released in Japan as Esteban, Child of the Sun (Japanese: 太陽の子エステバン, Hepburn: Taiyō no Ko Esuteban) and in France as Les Mystérieuses Cités d'or, is a French-Japanese animated series co-produced by DiC Audiovisuel and Studio Pierrot. Set in 1532, the series follows the adventures of a young Spanish boy named Esteban who joins a voyage to the New World in search of the lost Seven Cities of Gold and his father.The series was originally broadcast in Japan and the French version, edited to have different characterization and music, was subsequently redubbed and distributed in many different countries throughout the world. It is currently licensed for English language home video release in the United Kingdom, Australia, and North America by Fabulous Films.

Thomas Thirlby

Thomas Thirlby (or Thirleby; c. 1506–1570), was the first and only bishop of Westminster (1540–50), and afterwards successively bishop of Norwich (1550–54) and bishop of Ely (1554–59). While he acquiesced in the Henrician schism, with its rejection in principle of the Roman papacy, he remained otherwise loyal to the doctrine of the Roman Catholic Church during the English Reformation.

Tiger conservation

The tiger is an iconic species. Tiger conservation attempts to prevent the animal from becoming extinct and preserving its natural habitat. This is one of the main objectives of the international animal conservation community. The Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES) has played a crucial role in improving international efforts for tiger conservation.

Wildlife trade

Wildlife trade refers to the commerce of products that are derived from non-domesticated animals or plants usually extracted from their natural environment or raised under controlled conditions. It can involve the trade of living or dead individuals, tissues such as skins, bones or meat, or other products. Legal wildlife trade is regulated by the United Nations' Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES), which currently has 183 member countries called Parties. Illegal wildlife trade is widespread and constitutes one of the major illegal economic activities, comparable to the traffic of drugs and weapons. Wildlife trade is a serious conservation problem, has a negative effect on the viability of many wildlife populations and is one of the major threats to the survival of vertebrate species.

IUCN Red List
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