Bolboceratidae, the dor beetles, is a family of beetle.[1] It was historically treated as a subfamily of the earth-boring dung beetles (family Geotrupidae), but has been considered a separate family since 1995.[2] Some recent classifications have persisted in treating bolboceratids as a subfamily (e.g.[3][4]) but these classifications are contradicted by recent phylogenetic studies of relationships indicating that bolboceratids are not closely related to geotrupids (e.g., that bolboceratids are more closely related to Pleocomidae and Passalidae[5]).

Scientific classification

Mulsant, 1842



  • Athyreus
  • Australobolbus
  • Blackbolbus
  • Blackburnium
  • Bolbaffer
  • Bolbaffroides
  • Bolbelasmus
  • Bolbobaineus
  • Bolbocaffer
  • Bolbocerastes
  • Bolboceratex
  • Bolboceratops
  • Bolbocerodema
  • Bolboceroides
  • Bolbocerosoma
  • Bolbochromus
  • Bolbogonium
  • Bolbohamatum
  • Bolboleaus
  • Bolborhachium
  • Bolborhinum
  • Bolborhombus
  • Bolbothyreus
  • Bolbotrypes
  • Bradycinetulus
  • Cretobolbus
  • Elephastomus
  • Eubolbitus
  • Eucanthus
  • Gilletinus
  • Halffterobolbus
  • Meridiobolbus
  • Mesoathyreus
  • Mimobolbus
  • Namibiobolbus
  • Namibiotrupes
  • Neoathyreus
  • Odonteus
  • Parabolbapium
  • Parathyreus
  • Pereirabolbus
  • Prototrupes
  • Pseudoathyreus
  • Socotrabolbus
  • Somalobolbus
  • Senaspidius
  • Zefevazia


  1. ^ Gomes Gonçalves, Marcos Paulo (December 2017). "Relationship Between Meteorological Conditions and Beetles in Mata de Cocal". Revista Brasileira de Meteorologia. 32 (4): 543–554. doi:10.1590/0102-7786324003. ISSN 0102-7786.
  2. ^ Bolboceratidae at TOL
  3. ^ Smith, A.B.T. (2006) A Review of the Family-group Names for the Superfamily Scarabaeoidea (Coleoptera) with Corrections to Nomenclature and a Current Classification. The Coleopterists Bulletin, 60:144-204.
  4. ^ Bouchard, P.; Bousquet, Y.; Davies, A.; Alonso-Zarazaga, M.; Lawrence, J.; Lyal, C.; Newton, A.; Reid, C.; Schmitt, M.; Ślipiński, A.; Smith, A. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys. 88 (88): 1–972. doi:10.3897/zookeys.88.807. PMC 3088472. PMID 21594053.
  5. ^ MCKENNA, D. D., WILD, A. L., KANDA, K., BELLAMY, C. L., BEUTEL, R. G., CATERINO, M. S., FARNUM, C. W., HAWKS, D. C., IVIE, M. A., JAMESON, M. L., LESCHEN, R. A. B., MARVALDI, A. E., MCHUGH, J. V., NEWTON, A. F., ROBERTSON, J. A., THAYER, M. K., WHITING, M. F., LAWRENCE, J. F., ŚLIPIŃSKI, A., MADDISON, D. R. and FARRELL, B. D. (2015), The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Syst Entomol, 40: 835–880. doi:10.1111/syen.12132

External links


Boganiidae is a family of beetles, in the suborder Polyphaga.


Bolbelasmus is a genus of beetles in the family Geotrupidae. There are at least 20 described species in Bolbelasmus.


Bolbocerosoma (sometimes misspelled as Bolbocerasoma) is a genus of earth-boring scarab beetles in the family Bolboceratidae. There are about 13 described species in Bolbocerosoma.

Bolbocerosoma pusillum

Bolbocerosoma pusillum is a species of beetle in the family Bolboceratidae. It is found in North America.


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Geotrupidae (from Greek geos, earth, and trypetes, borer) is a family of beetles in the order Coleoptera. They are commonly called earth-boring dung beetles. Most excavate burrows in which to lay their eggs. They are typically detritivores, provisioning their nests with leaf litter (often moldy), but are occasionally coprophagous, similar to dung beetles. The eggs are laid in or upon the provision mass and buried, and the developing larvae feed upon the provisions. The burrows of some species can exceed 2 metres in depth.

A few species communicate by stridulation (rubbing body parts together to make sounds).


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Odonteus is a genus of beetles in the family Bolboceratidae, formerly known by the name Bolboceras. The name Bolboceras was replaced in 1990 when it was discovered that Odonteus had been published one month earlier; this finding was upheld by ICZN Opinion 2138 in 2006, which made the two names synonymous, with Odonteus having priority. Some researchers continue to use the invalid junior name (e.g.).

Odonteus obesus

Odonteus obesus is a species of earth-boring scarab beetle in the family Bolboceratidae. It is found in North America.


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Extant Coleoptera families


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