Allodaposuchidae

Allodaposuchidae is an extinct clade of basal eusuchians that lived in southern Europe during the Late Cretaceous (Santonian-Maastrichtian).[1][2]

Allodaposuchidae
Temporal range: Late Cretaceous, Santonian–Maastrichtian
Allodaposuchus precedens
A. precedens skull
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Eusuchia
Family: Allodaposuchidae
Narváez et al., 2015
Type species
Allodaposuchus precedens
Nopcsa, 1928
Genera

Agaresuchus
Allodaposuchus
Arenysuchus
Ischyrochampsa
Lohuecosuchus
Massaliasuchus
Musturzabalsuchus

Systematics

The type genus, Allodaposuchus, was originally described in 1928 from the Maastrichtian-age Sard Formation of the Hațeg Basin in Transylvania, Romania, and classified as a relative of the North American Leidyosuchus.[3] It was later classified as a eusuchian outside of Crocodylia in a 2001 paper, and subsequent studies found a number of European eusuchian species (Arenysuchus, Ischyrochampsa, Massaliasuchus, Musturzabalsuchus) to group with Allodaposuchus, prompting the erection of the clade Allodaposuchidae to accommodate Allodaposuchus and all European eusuchians closely related to it.[1][2][4][5][6][7]

References

  1. ^ a b Narváez, Iván; Brochu, Christopher A.; Escaso, Fernando; Pérez-García, Adán; Ortega, Francisco (2015). "New crocodyliforms from southwestern Europe and definition of a diverse clade of European Late Cretaceous basal eusuchians". PLOS ONE. 10 (11): e0140679. doi:10.1371/journal.pone.0140679. PMC 4633049. PMID 26535893.
  2. ^ a b Narváez, I.; Brochu, C.A.; Escaso, F.; Pérez-García, A.; Ortega, F. "New Spanish Late Cretaceous eusuchian reveals the synchronic and sympatric presence of two allodaposuchids". Cretaceous Research. 65: 112–125. doi:10.1016/j.cretres.2016.04.018.
  3. ^ Nopcsa F., 1928 – Paleontological notes on Reptilia. 7. Classification of the Crocodilia – Geologica Hungarica, Series Palaeontologica 1: 75–84.
  4. ^ Buscalioni, A. D.; Ortega, F.; Weishampel, D. B.; Jianu, C. M. (2001). "A revision of the crocodyliform Allodaposuchus precedens from the Upper Cretaceous of the Hateg Basin, Romania. Its relevance in the phylogeny of Eusuchia". Journal of Vertebrate Paleontology. 21: 74–86. doi:10.1671/0272-4634(2001)021[0074:AROTCA]2.0.CO;2.
  5. ^ Martin, J. E.; Delfino, M.; Garcia, G.; Godefroit, P.; Berton, S.; Valentin, X. (2016). "New specimens of Allodaposuchus precedens from France: intraspecific variability and the diversity of European Late Cretaceous eusuchians". Zoological Journal of the Linnean Society. 176: 607–631. doi:10.1111/zoj.12331.
  6. ^ Blanco, Alejandro; Fortuny, Josep; Vicente, Alba; Luján, Àngel H.; García-Marçà, Jordi Alexis; Sellés, Albert G. (2015). "A new species of Allodaposuchus (Eusuchia, Crocodylia) from the Maastrichtian (Late Cretaceous) of Spain: Phylogenetic and paleobiological implications". PeerJ. 3: e1171. doi:10.7717/peerj.1171. PMC 4558081. PMID 26339549.
  7. ^ Blanco, A.; Puértolas-Pascual, E.; Marmi, J.; Vila, B.; Sellés, A. G. (2014). "Allodaposuchus palustris sp. nov. From the Upper Cretaceous of Fumanya (South-Eastern Pyrenees, Iberian Peninsula): Systematics, Palaeoecology and Palaeobiogeography of the Enigmatic Allodaposuchian Crocodylians". PLoS ONE. 9 (12): e115837. doi:10.1371/journal.pone.0115837. PMC 4281157. PMID 25551630.
Aegyptosuchidae

Aegyptosuchidae is an extinct family of eusuchian crocodyliforms from the Cretaceous period of Africa. They are characterized by their large size and flat heads. The family includes two genera, Aegyptosuchus and Aegisuchus.

Agaresuchus

Agaresuchus is an extinct genus of allodaposuchid once known as "Allodaposuchus" subjuniperus.

In 2013 a second species of Allodaposuchus, A. subjuniperus, was named on the basis of a skull from the late-Maastrichtian Conquès Formation, part of the Tremp Group, in the province of Huesca, Spain. The skull was found underneath a juniper tree whose roots had grown between the bones, hence the species name subjuniperus or "under juniper" in Latin. In 2016, the species was renamed to Agaresuchus subjuniperus The 2013 study also proposed that the French and Spanish fossils assigned to A. precedens in 2001 might actually represent a new unnamed species of Allodaposuchus currently identified as Allodaposuchus sp.

Alligatorium

Alligatorium is an extinct genus of atoposaurid crocodylomorph from Late Jurassic marine deposits in France.

Amphicotylus

Amphicotylus is an extinct genus of goniopholidid mesoeucrocodylian from the Tithonian of Colorado and Oklahoma.

Anthracosuchus

Anthracosuchus (meaning "coal crocodile" in Greek) is an extinct genus of dyrosaurid crocodyliform from the Paleocene of Colombia. Remains of Anthracosuchus balrogus, the only known species, come from the Cerrejón Formation in the Cerrejón mine, and include four fossil specimens with partial skulls. Anthracosuchus differs from other dyrosaurids in having an extremely short (brevirostrine) snout, widely spaced eye sockets with bony protuberances around them, and osteoderms that are smooth and thick. It is one of the most basal dyrosaurids along with Chenanisuchus and Cerrejonisuchus. The species name is a reference to the Balrog, a creature in J. R. R. Tolkien's fantasy novel The Lord of the Rings that could, like the remains of Anthracosuchus, be found in a mine.

Atoposauridae

Atoposauridae is a family of crocodile-line archosaurs belonging to Neosuchia. The majority of the family are known from Late Jurassic to Early Cretaceous marine deposits in France, Portugal, and Bavaria in southern Germany. The discovery of the genus Aprosuchus, however, extends the duration of the lineage to the end of the Cretaceous in Romania.

Brillanceausuchus

Brillanceausuchus is an extinct genus of atoposaurid crocodylomorph. Fossils have been found in Early Cretaceous–age rocks of Cameroon. The genus is notable for the position of the secondary choana within its palate. Parts of the pterygoid bones make up the rostral margin of the choana and thus separate it from the palatines, a feature also seen in the more advanced neosuchian suborder Eusuchia. This characteristic was once thought to be characteristic of Eusuchia, but its presence in Brillanceausuchus suggests that the trait is homoplasic, thus making the evolution of the position of the choana within crocodilians more complex than previously thought.

Coelosuchus

Coelosuchus is an extinct genus of goniopholidid mesoeucrocodylian. Fossils have been found from the Graneros Shale of the Benton Group in Wyoming, and are of Cenomanian age. It was slightly over 1 meter in length.

Eusuchia

The Eusuchia ("true crocodiles") are a clade of crocodylomorphs that first appears in the Early Cretaceous with Hylaeochampsa. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survived the K-T extinction. Since the other two clades died out 35 and 11 million years ago, all living crocodilian species are eusuchians, as are many extinct forms.

Ischyrochampsa

Ischyrochampsa is an extinct genus of Late Cretaceous mesoeucrocodylian belonging to the eusuchian clade Allodaposuchidae. Fossils of the type species I. meridionalis are late Campanian in age and were found in the commune of Saint-Estève-Janson in Bouches-du-Rhône, France. Material is also known from Spain. At an estimated length of over 4 metres (13 ft), Ischyrochampsa was a large mesoeucrocodylian. It was named and described in 1995.Ischyrochampsa was first classified as a trematochampsid, but was removed from the group by subsequent studies. In their description of Allodaposuchus remains from southern France, Martin and his colleagues treated the genus as a possible junior synonym of Allodaposuchus.

Karatausuchus

Karatausuchus is an extinct genus of atoposaurid crocodylomorph. It is known from a single specimen discovered in the Late Jurassic (Oxfordian - Kimmeridgian) Karabastau Svita from the vicinity of Mikhailovka in the Karatau Mountains of southern Kazakhstan. The type specimen is PIN 25858/1, a complete but poorly preserved juvenile skeleton with some possible soft tissue preservation. It is notable for having over 90 teeth, but its other anatomical details are difficult to discern. The length of this individual is estimated at 160 millimetres (6.3 in). Karatausuchus was described in 1976 by Mikhail Efimov, and the type species is K. sharovi.

Khoratosuchus

Khoratosuchus is an extinct genus of neosuchian crocodylomorph which existed in northeast Thailand during the early Cretaceous period. Its type species is Khoratosuchus jintasakuli. Khoratosuchus is the youngest and most advanced Mesozoic crocodyliform yet known from Thailand. It possesses several distinctive features that help determine its phylogenetic position among crocodylomorphs, including secondary choanae relatively posterior and almost encircled by the pterygoid bones on the palate and a smooth dorsal surface of the skull.

Laganosuchus

Laganosuchus is an extinct genus of stomatosuchid crocodyliform. Fossils have been found from Niger and Morocco and date back to the Upper Cretaceous.

Massaliasuchus

Massaliasuchus is an extinct genus of allodaposuchid crocodylian that is known from fossils found in Santonian–Campanian-age Upper Cretaceous rocks of southeastern France.

Musturzabalsuchus

Musturzabalsuchus is an extinct genus of crocodylian, and one of the oldest known members of the superfamily Alligatoroidea. The generic name means "Broadened rostrum crocodile", with "Musturzabal" meaning "broadened rostrum" in Basque and "suchus" meaning "crocodile" in Greek. The type and only species is M. buffetauti, named after the French paleoherpetologist Eric Buffetaut. The material first assigned to Musturzabalsuchus in 1997 has been found from the locality of Laño in Condado de Treviño, northern Spain. Although dating back to the Late Cretaceous, the exact age of the strata in which material of Musturzabalsuchus occurs in the locality is not known: it is either Late Campanian or very Early Maastrichtian. Despite the unusually high quantity of remains belonging to the genus (most other continental Late Cretaceous crocodilian genera from Europe are poorly represented), the only skeletal elements known from Musturzabalsuchus are the maxilla and mandible. Some fragments of these bones have been found from the locality of Armuña in the province of Segovia that were previously referred to an unnamed trematochampsid. Like the holotype and paratype material found from Laño, these fossils, known collectively as UPUAM-502, are Campano-Maastrichtian in age. Another specimen (MHNM 10834.0) from the Fuvelian Lignites of France has been referred to Musturzabalsuchus in 1999. However, the characteristics used to assign the material to better-known specimens of Musturzabalsuchus from Spain were questioned in a later study. Material from Musturzabalsuchus has been found more recently from Valencia, Spain, being slightly older in age than specimens from other localities, dating back to the Early or Middle Campanian.

Nannosuchus

Nannosuchus (meaning "dwarf crocodile") is an extinct genus of goniopholidid mesoeucrocodylian from the Berriasian of England.

Symptosuchus

Symptosuchus is an extinct genus of goniopholidid mesoeucrocodylian. It is known from the Late Cretaceous of Argentina. Argentine paleontologist Florentino Ameghino named the genus in 1899, along with the type species S. contortidens. It was formally described by Carlos Rusconi in 1934.

Wahasuchus

Wahasuchus is a genus of extinct mesoeucrocodylian of the Middle Campanian age found in the Quseir Formation, Egypt. The generic name derives from the Arabic word واحة (waha), which means "oasis", and souchos from the Greek in honor of crocodile-headed god of ancient Egypt. The specific egyptensis (Lat.) means from Egypt.Fossils of skull and jaw fragments, dorsal vertebrae, and fragmentary appendicular remains have been recovered.

Extinct crocodilian species

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