The Aderidae, sometimes called ant-like leaf beetles, are a family of beetles that bear some resemblance to ants. The family consists of about 1,000 species in about 50 genera, of which most are tropical, although overall distribution is worldwide.

As with the Anthicidae, their heads constrict just in front of the pronotum, forming a neck, although the posterior end of the pronotum is not usually as narrow. The eyes are hairy with a granular appearance. The first two abdominal sternites are fused, and in only some groups is a suture even visible. Sizes are 1–4 mm.

As the name suggests, most adults are found on the undersides of the leaves of shrubs and trees, while larvae have found in rotting wood, leaf litter, and nests of other insects.

As of 2002, the last publication of a world catalog of the family was that of Maurice Pic in 1910.

Synonyms of the family include:

  • Xylophilidae Shuckard 1840
  • Euglenesidae or Euglenidae Seidlitz 1875
  • Hylophilidae Pic 1900
Scraptogetus cf.anthracinus
Scraptogetus cf. anthracinus
Scientific classification

Winkler, 1927

about 50 genera




Aderus is a genus of leaf beetles of the family Aderidae that resemble ants. The genus was named by John Obadiah Westwood in 1829.

Aderus populneus

Aderus populneus is a species of ant-like leaf beetle in the family Aderidae. Native to the west Palaearctic (very common in England, it has been introduced in North America.. Large eyes, pubescent antennae, 1.5-2mm


Ariotus is a genus of ant-like leaf beetles in the family Aderidae. There are at least four described species in Ariotus.

Ariotus subtropicus

Ariotus subtropicus is a species of ant-like leaf beetle in the family Aderidae. It is found in North America.


Cnopus is a genus of ant-like leaf beetles in the family Aderidae. There are at least four described species in Cnopus.


Elonus is a genus of ant-like leaf beetles in the family Aderidae. There are about seven described species in Elonus.


Emelinus is a genus of ant-like leaf beetles in the family Aderidae. There are at least three described species in Emelinus.


Euglenes is a genus of ant-like leaf beetles in the family Aderidae. There are about five described species in Euglenes.


Ganascus is a genus of ant-like leaf beetles in the family Aderidae. There are at least two described species in Ganascus.


Gymnoganascus is a genus of ant-like leaf beetles in the family Aderidae. There is one described species in Gymnoganascus, G. stephani. The distribution range of Gymnoganascus stephani includes the Caribbean, Central America, and North America.

List of fauna of Batu Caves

The following species have been collected from the Dark Cave, Batu Caves, Selangor in Malaysia:

Pheretima indica (Haplotaxida: Magascolecidae)

Dichogaster sp. (Haplotaxida: Octochaetidae[1])

Dichogaster bolavi (Haplotaxida: Octochaetidae)

Sarax brachydactylus (Amblypygi: Tarantulidae)

Damarchus cavernicolus (Araneae: Ctenizidae)

Heteropoda robusta (Araneae: Heteropodidae)

Liphistius batuensis (Araneae: Liphistiidae)

Psiluderces crinitus (Araneae: Ochyroceratidae)

Spermophora miser (Araneae: Pholcidae)

Psechrus curvipalpus (Araneae: Psechridae)

Scytodes magnus (Araneae: Scytodidae)

Theriheridion rufipes (Araneae: Therididae)

Uloborus spelaeus (Araneae: Uloboridae)

Paracheyletia sp. (Eleutherengona: Cheyletidae)

Ornithodorus bauensis (Ixodida: Argasidae)

Belba sp. (Oribatida: Belbidae)

Galumna sp. (Oribatida: Galumnidae)

Vaghia sp. (Oribatida: Galumnidae)

Scheloribates exuvium (Oribatida: Oribatulidae)

Scheloribates sp. (Oribatida: Oribatulidae)

Trombicula batui (Parasitengona: Trombiculidae)

Trombicula insolli (Parasitengona: Trombiculidae)

Cryptocheiridium lucifugum (Pseudoscorpionida: Ideronicidae)

Dhanus sumatranus (Pseudoscorpionida: Ideronicidae)

Dhanus doveri (Pseudoscorpionida: Ideronicidae)

Cunaxa setirostris (Trombidiformes: Cunaxidae)

Scutigera decipiens (Scutigeromorpha: Cambalidae)

Glyphiulus sp. (Sphaerotheriida: Cambalidae)

Doratodesmus sp. (Sphaerotheriida: Cambalidae)

Brachystomella contorta (Collembola: Brachystomellidae)

Cunaxa spp. (Collembola: Cunaxidae)

Willemia nadchatrami (Collembola: Neogastruridae)

Aderus mcclurei (Coleoptera: Aderidae)

Euglenes batuensis (Coleoptera: Aderidae)

Euglenes cephalicus (Coleoptera: Aderidae)

Euglenes malayanus (Coleoptera: Aderidae)

Euglenes troglodytes (Coleoptera: Aderidae)

Unidentified (Coleoptera: Carabidae)

Unidentified (Coleoptera: Curculionidae)

Aethriostoma undulata (Coleoptera: Dermestidae)

Trinodes sp. (Coleoptera: Dermestidae)

Cardiophorus carduelis (Coleoptera: Elateridae)

Melanoxanthus near dohrni (Coleoptera: Elateridae)

Platynuchus sp. (Coleoptera: Elateridae)

Unidentified (Coleoptera: Endomychidae)

Platycladoxena near angulosa (Coleoptera: Erotylidae)

Thallisellodes limbooliati (Coleoptera: Erotylidae)

Cercyon gebieni (Coleoptera: Hydrophilidae)

Dactylosternum abdominale (Coleoptera: Hydrophilidae)

Lychnocrepis antricola (Coleoptera: Lampyridae)

Unidentified (Coleoptera: Melyridae)

Unidentified (Coleoptera: Nitidulidae)

Unidentified (Coleoptera: Orthoperidae)

Unidentified (Coleoptera: Pselaphidae)

Trox costatus (Coleoptera: Scarabaeidae)

Unidentified (Coleoptera: Scydmaenidae)

Unidentified (Coleoptera: Silphidae)

Unidentified (Coleoptera: Staphylinidae)

Coeloecetes cavernicola (Coleoptera: Tenebrionidae)

Chelisoches morio (Dermaptera: Forficulidae)

Chelisoches brevipennis (Dermaptera: Forficulidae)

Unidentified (Diptera: Cecidomyidae)

Atrichopogon jacobsoni (Diptera: Ceratopogonidae)

Culicoides huffi (Diptera: Ceratopogonidae)

Culicoides peregrinus (Diptera: Ceratopogonidae)

Culicoides prob. arakawai (Diptera: Ceratopogonidae)

Forcipomyia spp. (Diptera: Ceratopogonidae)

Stilobezzia sp. (Diptera: Ceratopogonidae)

Eusmittia cavernae (Diptera: Chironomidae)

Paratendipes inarmatus (Diptera: Chironomidae)

Pentaneura batuensis (Diptera: Chironomidae)

Podonomus sp. (Diptera: Chironomidae)

Tricimba batucola (Diptera: Chloropidae)

Chyromya prob. dubia (Diptera: Chyromyidae)

Gymnochiromyia sp. (Diptera: Chyromyidae)

Aedes albopictus (Diptera: Culicidae)

Culex tritaeniorhynchus (Diptera: Culicidae)

Uranotaenia sp. (Diptera: Culicidae)

Condystylus sp. (Diptera: Dolichopodidae)

Drosophila ananassae (Diptera: Drosophilidae)

Drosophila melanogaster (Diptera: Drosophilidae)

Desmometopa spp. (Diptera: Milichiidae)

Leptometopa mcclurei (Diptera: Milichiidae)

Milichia sp. (Diptera: Milichiidae)

Phyllomyza cavernae (Diptera: Milichiidae)

Fannia leucosticta (Diptera: Muscidae)

Ophyra chalcogaster (Diptera: Muscidae)

Chetoneura cavernae (Diptera: Mycetophilidae)

Eucampsipoda sundaicum (Diptera: Nycteribiidae)

Nycteribosca prob. gigantea (Diptera: Nycteribiidae)

Diploneura peregrina (Diptera: Phoridae)

Brunettia sp. (Diptera: Psychodidae)

Phlebotomus anodontis (Diptera: Psychodidae)

Phlebotomus anondontis (Diptera: Psychodidae)

Phlebotomus argentipes (Diptera: Psychodidae)

Phlebotomus asperules (Diptera: Psychodidae)

Phlebotomus lutea (Diptera: Psychodidae)

Phlebotomus makati (Diptera: Psychodidae)

Phlebotomus malayica (Diptera: Psychodidae)

Phlebotomus savaiiensis (Diptera: Psychodidae)

Phlebotomus stantion (Diptera: Psychodidae)

Psychoda acanthostyla (Diptera: Psychodidae)

Psychoda alternata (Diptera: Psychodidae)

Psychoda aponesos (Diptera: Psychodidae)

Psychoda harrisi (Diptera: Psychodidae)

Psychoda malleola (Diptera: Psychodidae)

Psychoda pellucida (Diptera: Psychodidae)

Psychoda vagabunda (Diptera: Psychodidae)

Sycorax malayensis (Diptera: Psychodidae)

Telmatoscopus albipuntatus (Diptera: Psychodidae)

Telmatoscopus kulas (Diptera: Psychodidae)

Telmatoscopus mcclurei (Diptera: Psychodidae)

Trichomyia batu (Diptera: Psychodidae)

Trichomyia malaya (Diptera: Psychodidae)

Unidentified (Diptera: Sarcophagidae)

Bradysia flagellicornis (Diptera: Sciaridae)

Bradysia leucocerca (Diptera: Sciaridae)

Bradysia platytergum (Diptera: Sciaridae)

Bradysia spp. (Diptera: Sciaridae)

Bradysia leucocerca (Diptera: Sciaridae)

Phorodonta malayana (Diptera: Sciaridae)

Plastosciara near brevicalcarata (Diptera: Sciaridae)

Soudekia sp. (Diptera: Sciaridae)

Leptocera brevicostata (Diptera: Sphaeroceridae)

Sargus metallinus (Diptera: Stratiomyidae)

Helius cavernicolus (Diptera: Tipulidae)

Cleon sp. (Ephemeroptera: Ephemeridae)

Myiophanes fluitaria (Hemiptera: Reduviidae)

Unidentified (Hemiptera: Cydnidae)

Bagauda lucifigus (Hemiptera: Reduviidae)

Reduvius gua (Hemiptera: Reduviidae)

Fulvinus brevicornis (Hemiptera: Miridae)

Apanteles carpatus (Hymenoptera: Braconidae)

Apanteles sp. (Hymenoptera: Braconidae)

Aulosaphes sp. (Hymenoptera: Braconidae)

Epitranus lacteipennis (Hymenoptera: Chalcididae)

Epitranus stantoni (Hymenoptera: Chalcididae)

Bathroponera rufipes (Hymenoptera: Formicidae)

Bathroponera tridentata (Hymenoptera: Formicidae)

Camponotus sp. (Hymenoptera: Formicidae)

Leptogenys diminuta (Hymenoptera: Formicidae)

Monomorium pharaonis (Hymenoptera: Formicidae)

Paratrechina longicornis (Hymenoptera: Formicidae)

Pheidole javana (Hymenoptera: Formicidae)

Ponera sp. (Hymenoptera: Formicidae)

Pristomyrmex sp. (Hymenoptera: Formicidae)

Tapinoma melanocephalum (Hymenoptera: Formicidae)

Hypsicerca cavicola (Hymenoptera: Ichneumonidae)

Hypsicerca fullawayi (Hymenoptera: Ichneumonidae)

Trogaspidio sp. (Hymenoptera: Mutillidae)

Opogona cerodelta (Lepidoptera: Lyonetidae)

Attacus atlas (Lepidoptera: Saturnidae)

Tinea antricola (Lepidoptera: Tineidae)

Tinea palaechrysis (Lepidoptera: Tineidae)

Neglurus vitripennis (Neuroptera: Myrmeleonidae)

Diestrammena gravely (Orthoptera: Gryllacridae)

Gryllotalpa fulvipes (Orthoptera: Gryllotalpidae)

Mermecophilus dubius (Orthoptera: Gryllidae)

Pycnoscelus striatus (Orthoptera: Blattidae)

Liposcelis sp. (Psocoptera: Liposcelidae)

Parasoa haploneura (Psocoptera: Lepidopsocidae)

Psyllipsocus batuensis (Psocoptera: Psyllipsocidae)

Ectopsocus maindroni (Psocoptera: Peripsocidae)

Parasoa haploneura (Psocoptera: Forficulidae)

Thaumapsylla breviceps orientalis (Siphonaptera: Ischnopsyllidae)

Parabathynella malaya (Bathynellacea: Bathynellidae)

Armadillo intermixtus (Isopoda: Armadillidiidae)

Philoscia dobakholi (Isopoda: Oniscidae)

Unidentified (Podocopida: Cyprididae)

Chaerilus prob. celebensis (Scorpionida: Chactidae)

Bufo asper (Anura: Bufonidae)

Bufo melanostictus (Anura: Bufonidae)

Callula pulchra (Anura: Microhylidae)

Rana calconota (Squamata: Scincidae)

Collocalia sp. (Apodiformes: Apodidae)

Hirundo daurica (Passeriformes: Hirundinidae)

Myophoneus flavirostris (Passeriformes: Turdidae)

Taphozous melanopogon (Chiroptera: Emballonuridae)

Eonycteris spelaea (Chiroptera: Pteropodidae)

Hipposideros armiger debilis (Chiroptera: Rhinolophidae)

Hipposideros bicolor (Chiroptera: Rhinolophidae)

Hipposideros diadema vicarius (Chiroptera: Rhinolophidae)

Hipposideros galiterus (Chiroptera: Rhinolophidae)

Rhinolophus affinis superans (Chiroptera: Rhinolophidae)

Rhinolophus luctus morio (Chiroptera: Rhinolophidae)

Myotis mystacinus (Chiroptera: Vespertilionidae)

Rattus rattus jalorensis (Rodentia: Muridae)

Crocidura malayana (Soricomorpha: Soricidae)

Elaphe taeniura (Squamata: Colubridae)

Dryophiops rubescens (Squamata: Colubridae)

Ahaetulla formosa (Squamata: Colubridae)

Gecko marmorata (Squamata: Gekkonidae)

Lygosoma scotophilum (Squamata: Scincidae)

Paludomus buccula, Paludomus buccula var. minuta (Thiaridae)

Opeas doveri (Stylommatophora: Stenogyridae)

Opeas dimorpha (Stylommatophora: Stenogyridae)

Achatina fulica (Pulmonata: Achatinidae)

Dugesia sp. (Seriata: Scincidae)


Pseudariotus is a genus of ant-like leaf beetles in the family Aderidae. There is at least one described species in Pseudariotus, P. notatus.


The family Pythidae is a small group of beetles with no vernacular common name, though recent authors have coined the name dead log bark beetles. There are about 10 genera and 17 described species in Pythidae.

Qadrius salvus

Qadrius salvus is a beetle species in the Aderidae family.


The Tenebrionoidea are a very large and diverse superfamily of beetles.

It generally corresponds to the Heteromera of earlier authors.

It contains these families:

Aderidae Winkler 1927 (ant-like leaf beetles)

Anthicidae Latreille 1819 (ant-like flower beetles)

Archeocrypticidae Kaszab 1964

Boridae C. G. Thomson 1859

Chalcodryidae Watt 1974

Ciidae Leach 1819 (minute tree-fungus beetles) (= Cisidae)

Melandryidae Leach 1815 (false darkling beetles)

Meloidae Gyllenhal 1810 (blister beetles)

Mordellidae Latreille 1802 (tumbling flower beetles)

Mycetophagidae Leach 1815 (hairy fungus beetles)

Mycteridae Blanchard 1845

Oedemeridae Latreille 1810 (false blister beetles)

Perimylopidae St. George 1939

Prostomidae C. G. Thomson 1859

Pterogeniidae Crowson 1953

Pyrochroidae Latreille 1807 (fire-colored beetles, etc.)

Pythidae Solier 1834

Ripiphoridae Gemminger and Harold 1870 (wedge-shaped beetles) (= Rhipiphoridae)

Salpingidae Leach 1815 (narrow-waisted bark beetles, etc.)

Scraptiidae Mulsant 1856 (false flower beetles)

Stenotrachelidae C. G. Thomson 1859 (false long-horned beetles) (= Cephaloidae)

Synchroidae Lacordaire 1859

Tenebrionidae Latreille 1802 (darkling beetles)

Tetratomidae Billberg 1820

Trachelostenidae Lacordaire 1859

Trictenotomidae Blanchard 1845

Ulodidae Pascoe 1869

Zopheridae Solier 1834 (ironclad beetles, cylindrical bark beetles, etc.)


Vanonus is a genus of ant-like leaf beetles in the family Aderidae. There are about 16 described species in Vanonus.


Zonantes is a genus of ant-like leaf beetles in the family Aderidae. There are about 10 described species in Zonantes.

Zonantes hubbardi

Zonantes hubbardi is a species of ant-like leaf beetle in the family Aderidae. It is found in North America.

Zonantes nubifer

Zonantes nubifer is a species of ant-like leaf beetle in the family Aderidae. It is found in North America.

Extant Coleoptera families

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